Family HESPERIIDAE Latreille, 1809 – SKIPPERS

  Subfamily Eudaminae

Phocides  Hübner, [1819]

Phocides polybius (Fabricius, 1793)  Guava Skipper  TL: Indiis

Phocides polybius lilea (Reakirt, [1867])  S Texas to Costa Rica  M T S1 S2 L I1 I2 G

Phocides polybius polybius (Fabricius, 1793)  Panama, Colombia, Venezuela, Guianas, Trinidad  M T S L G

Phocides polybius phanias (Burmeister, 1880)  TL: Argentina  M S

Phocides charon (C. Felder & R. Felder, 1859)  TL: Brazil  M T S IL

Phocides distans (Herrich-Schäffer, 1869)  TL: ?

Phocides distans licinus (Möschler, 1879)  Honduras, Costa Rica, Panama  M T S L G

Phocides distans distans (Herrich-Schäffer, 1869)  TL: ?  M T S IL

Phocides distans silva Evans, 1952  TL: Peru  M T S

Phocides perillus (Mabille, 1888)  TL: Colombia  M T OI S

Phocides metrodorus E. Bell, 1932  Metrodorus Skipper  TL: Colombia

Phocides metrodorus nigrescens E. Bell, 1938  Costa Rica, Colombia, Venezuela, Ecuador, Peru; species?  M T S1 S2 I1 I2

Phocides metrodorus metrodorus E. Bell, 1932  Panama to Bolivia and Brazil  M T S G

Phocides metrodorus metron Evans, 1952  TL: Paraguay  M T OD G

Phocides vulcanides Röber, 1925  TL: Colombia  M T S

Phocides novalis Evans, 1952  TL: Peru  M T S G

Phocides thermus (Mabille, 1883)  Thermus Skipper  TL: Colombia

Phocides thermus thermus (Mabille, 1883)  SE Mexico to Venezuela and Ecuador  M T S G

Phocides thermus bellina Evans, 1952  TL: Ecuador  M T S

Phocides thermus valgus (Mabille, 1883)  TL: French Guiana  M T S

Phocides partia Evans, 1952  TL: Peru  M T S G

Phocides padrona Evans, 1952  TL: Peru  M T S G

Phocides belus Godman & Salvin, 1893  Belus Skipper  S Texas (one-time stray) to Costa Rica  M T S1 S2 L I

Phocides pigmalion (Cramer, 1779)  Pigmalion Skipper  TL: Surinam

Phocides pigmalion pigmalion (Cramer, 1779)  TL: Surinam  M T OI G

Phocides pigmalion hewitsonius (Mabille, 1883)  TL: Brazil (Am)  M T TI IL

Phocides pigmalion okeechobee (Worthington, 1881)  S Florida; mangroves  M T S L1 L2 I G

Phocides pigmalion batabanoides (W. Holland, 1902)  most Bahamian Islands  M T S

Phocides pigmalion batabano (Lucas, 1857)  Cuba, I. de Juventud, Andros, Little Cayman  M T S G

Phocides pigmalion bicolora (Boddaert, 1783)  Hispaniola  M T S

Phocides johnsoni E. Bell, 1947  TL: Colombia  M T S

Phocides urania (Westwood, 1852)  Urania Skipper  TL: Mexico ([Oax])

Phocides urania urania (Westwood, 1852)  Mexico, Guatemala; usually in montane forest  M T S L G

Phocides urania vida (A. Butler, 1872)  Costa Rica  M T S

Phocides pialia Hewitson, 1857  TL: Brazil

Phocides pialia maximus (Mabille, 1888)  TL: Brazil  M T OI S

Phocides pialia intermedia O. Mielke, 1992  TL: Brazil (MG)  M OI S

Phocides pialia pialia Hewitson, 1857  TL: Brazil  M T OI S G

Phocides oreides (Hewitson, [1875])  TL: Peru

Phocides oreides colombiana E. Bell, 1938  TL: Colombia  M T

Phocides oreides oreides (Hewitson, [1875])  TL: Peru  M T OI S IL

Phocides yokhara (A. Butler, 1870)  TL: Peru

Phocides yokhara yokhara (A. Butler, 1870)  TL: Peru  M T S

Phocides yokhara inca Le Cerf, 1922  TL: Peru  M T

Phocides yokhara dryas Le Cerf, 1922  TL: Peru  M T

Phocides yokhara charonotis (Hewitson, 1874)  TL: Bolivia  M T TI

Phocides lincea (Herrich-Schäffer, 1869)  TL: ?

Phocides lincea lincea (Herrich-Schäffer, 1869)  TL: ?  M S

Phocides lincea perkinsi (Kaye, 1931)  Jamaica  M T S

Hypocryptothrix  E. Watson, 1893

Hypocryptothrix teutas (Hewitson, 1876)  TL: Brazil (Am)  M T S

Tarsoctenus  E. Watson, 1893

Tarsoctenus papias Hewitson, 1857  TL: Brazil (Am)  M T OI S L

Tarsoctenus corytus (Cramer, 1777)  TL: Surinam

Tarsoctenus corytus gaudialis (Hewitson, 1876)  Nicaragua to Colombia  M T S L G

Tarsoctenus corytus corytus (Cramer, 1777)  TL: Surinam  M T OI OD S

Tarsoctenus corytus corba Evans, 1952  TL: Peru  M T S

Tarsoctenus praecia Hewitson, 1857  TL: Brazil (Pa)

Tarsoctenus praecia praecia Hewitson, 1857  TL: Brazil (Pa)  M T OI S

Tarsoctenus praecia rufibasis Boullet, 1910  TL: French Guiana  M T IL

Tarsoctenus praecia plutia Hewitson, 1857  TL: Brazil (Am)  M T OI S L

Tarsoctenus praecia luna Evans, 1952  TL: Bolivia  M T

Phanus  Hübner, [1819]

Phanus australis L. Miller, 1965  TL: Brazil (SC)  M T S

Phanus vitreus (Stoll, 1781)  Widespread Phanus  Oaxaca and Chiapas to southern Brazil  M T S1 S2 L I G

Phanus ecitonorum Austin, 1993  Army-ant Phanus  Costa Rica to southern Brazil  M S L

Phanus confusis Austin, 1993  Confusing Phanus  eastern Mexico to El Salvador  M T S

Phanus albiapicalis Austin, 1993  White-tipped Phanus  E & W Mexico to Panama, W Ecuador and Peru  M T S L

Phanus rilma Evans, 1952  West-Mexican Phanus  Pacific coast, Nayarit to Guatemala  M S G

Phanus grandis Austin, 1993  Grand Phanus  Nicaragua, Costa Rica, Panama, Colombia, Venezuela  M T S L

Phanus obscurior Kaye, 1925  Dark Phanus  TL: Trinidad

Phanus obscurior obscurior Kaye, 1925  SE Mexico to Colombia and Trinidad  M T S1 S2 I G

Phanus obscurior prestoni L. Miller, 1965  TL: Brazil (Am)  M T S

Phanus marshalli (W. F. Kirby, 1880)  Common Phanus  E & W Mexico to Bolivia  M S1 S2 L I G

Udranomia  A. Butler, 1870

Udranomia orcinus (C. Felder & R. Felder, 1867)  Orcinus Skipper  SE Mexico to Bolivia  M T S1 S2 IL I G

Udranomia kikkawai (Weeks, 1906)  Nervous Skipper  E & W Mexico to Brazil  M S1 S2 L I1 I2

Udranomia eurus (Mabille & Boullet, 1919)  Eurus Skipper  Costa Rica to Venezuela & Amazon basin  M S I

Udranomia spitzi (Hayward, 1942)  TL: Brazil (MT)  M S

Drephalys  E. Watson, 1893

Drephalys alcmon (Cramer, 1780)  Alcmon Skipper  Guatemala to Guianas and Brazil  M T S1 S2 S3 IL L I G

Drephalys mourei O. Mielke, 1968  TL: Brazil (SC)  M IL

Drephalys helixus (Hewitson, 1877)  Helixus Skipper  Panama  M T OD S IL G

Drephalys kidonoi Burns, 2000  Kidono's Skipper  W Costa Rica  M T S1 S2 I G

Drephalys phoenicoides (Mabille & Boullet, 1919)  Curious Skipper  Costa Rica to Guianas and NE Brazil  M S

Drephalys phoenice (Hewitson, 1867)  TL: Brazil (Am)  M T S IL

Drephalys heraclides E. Bell, 1942  TL: Peru  M T S

Drephalys miersi O. Mielke, 1968  TL: Brazil (SC)  M S IL

Drephalys opifex Evans, 1952  TL: Surinam  M OD

Drephalys olvina Evans, 1952  TL: Guyana  M S

Drephalys olva Evans, 1952  TL: Brazil (Pa)  M OD

Drephalys eous (Hewitson, 1867)  TL: Brazil (Pa)  M T S

Drephalys croceus Austin, 1995  TL: Brazil (Rd)  M OI G

Drephalys dumeril (Latreille, [1824])  Dumeril Skipper  Mexico to Ecuador and Brazil  M T OI OD S1 S2 IL G

Drephalys oria Evans, 1952  Oria Skipper  Mexico to Honduras  M T S G

Drephalys oriander (Hewitson, 1867)  Oriander Skipper  Mexico, Costa Rica to Guianas and N Brazil  M T S1 S2 IL G

Drephalys talboti (Le Cerf, 1922)  TL: French Guiana  M T S G

Drephalys tortus Austin, 1995  TL: Brazil (Rd)  M OI G

Augiades  Hübner, [1819]

Augiades crinisus (Cramer, 1780)  Crinisus Skipper  Costa Rica to Bolivia and Brazil  M T S L

Augiades epimethea (Plötz, 1883)  TL: Brazil

Augiades epimethea bicolor (Mabille & Boullet, 1919)  Costa Rica to Bolivia, Guianas and N Brazil  M T S

Augiades epimethea epimethea (Plötz, 1883)  TL: Brazil  M T

Hyalothyrus  Mabille, 1878

Hyalothyrus infernalis (Möschler, 1877)  TL: French Guiana; Surinam

Hyalothyrus infernalis infernalis (Möschler, 1877)  TL: French Guiana; Surinam  M T S

Hyalothyrus infernalis infa Evans, 1952  TL: Colombia  M T S

Hyalothyrus leucomelas (Geyer, 1832)  Leucomelas Skipper  Panama to Brazil and Peru  M S

Hyalothyrus nitocris (Stoll, 1782)  TL: Surinam  M S

Hyalothyrus mimicus Mabille & Boullet, 1919  TL: Peru  M S

Hyalothyrus neleus (Linnaeus, 1758)  TL: Indiis

Hyalothyrus neleus pemphigargyra (Mabille, 1888)  SE Mexico to Colombia and Venezuela  M T S L I G

Hyalothyrus neleus neleus (Linnaeus, 1758)  TL: Indiis  M T S L

Hyalothyrus neleus neda Evans, 1952  TL: Bolivia  M T S

Phareas  Westwood, 1852

Phareas coeleste Westwood, 1852  Magnificent Skipper  Costa Rica to Bolivia and Brazil  M T OI OD S1 S2 S3 S4 IL L I G

Entheus  Hübner, [1819]

Entheus eumelus (Cramer, 1777)  TL: Surinam  M T OI OD S IL G

Entheus ninyas H. Druce, 1912  TL: Bolivia  M T OI OD S

Entheus eunyas Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M OI S G

Entheus lemna (A. Butler, 1870)  TL: ?  M T OI OD S IL

Entheus gentius (Cramer, 1777)  TL: Surinam  M T OI OD TI S IL G Txt

Entheus bombus Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M OI S G

Entheus aureolus Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M OI S G

Entheus telemus Mabille, 1898  TL: Brazil  M OD IL G

Entheus latebrosus Austin, 1997  TL: Ecuador  M T OI

Entheus priassus (Linnaeus, 1758)  TL: Indiis

Entheus priassus priassus (Linnaeus, 1758)  TL: Indiis  M OI OD HS IL G Txt

Entheus priassus pralina Evans, 1952  TL: Brazil (ES)  M T IL G

Entheus aureanota Austin, O. Mielke & Steinhauser, 1997  TL: Brazil (Rd)  M OI G

Entheus curvus Austin, 1997  TL: Peru  M T OI G

Entheus matho Godman & Salvin, 1879  Giant Entheus  TL: Guatemala; Nicaragua

Entheus matho matho Godman & Salvin, 1879  SE Mexico to Costa Rica  M T OD S IL L G Txt

Entheus matho latifascius M. Hering, 1925  TL: Colombia  M OI OD S

Entheus matho aequatorius Mabille & Boullet, 1919  Costa Rica to Ecuador  M OD S IL G

Entheus matho dius Mabille, 1898  Panama Guianas, Peru, and Bolivia  M T OD TI S IL G

Entheus crux Steinhauser, 1989  Mexican Entheus  SE Mexico to Nicaragua  M T OD S G

Entheus warreni Grishin, 2012  Warren's Entheus  Ecuador  M T S G

Cabirus  Hübner, [1819]

Cabirus procas (Cramer, 1777)  TL: Surinam

Cabirus procas procas (Cramer, 1777)  TL: Surinam  M OI OD S IL

Cabirus procas junta Evans, 1952  TL: Peru  M T S1 S2 IL

Cabirus procas purda Evans, 1952  TL: Peru  M T S L

Proteides  Hübner, [1819]

Proteides mercurius (Fabricius, 1787)  Mercurial Skipper  TL: French Guiana

Proteides mercurius mercurius (Fabricius, 1787)  continental, S Texas to Argentina, SE Arizona as rare stray  M OI S1 S2 L I1 I2 G

Proteides mercurius sanantonio (Lucas, 1857)  Cuba, [one-time stray to Florida]  M S L

Proteides mercurius sanchesi E. Bell & W. Comstock, 1948  Hispaniola  M T S

Proteides mercurius jamaicensis Skinner, 1920  Jamaica  M T S

Proteides mercurius pedro (Dewitz, 1877)  Puerto Rico  M S

Proteides mercurius vincenti E. Bell & W. Comstock, 1948  St. Vincent  M T S

Proteides mercurius angasi Godman & Salvin, 1884  Dominica, St. Lucia, Tobago  M T S

Proteides mercurius grenadensis Enrico & Pinchon, 1969  Grenada  M S

Proteides maysi (Lucas, 1857)  Mays' Skipper  Cuba and Isla de Juventud  M T S

Epargyreus  Hübner, [1819]

Epargyreus zestos (Geyer, 1832)  Zestos Skipper  TL: "Surinam"

Epargyreus zestos zestos (Geyer, 1832)  Florida, Bahamas, Puerto Rico, Virgin Islands to St. Vincent  M T S L G

Epargyreus zestos inaguarum Clench & Bjorndal, 1980  Great Inagua, Crooked I., Turks and Caicos Islands  M T S

Epargyreus clarus (Cramer, 1775)  Silver-spotted Skipper  TL: [USA (Va)]

Epargyreus clarus californicus MacNeill, 1975  BC to Baja California Norte; Sierra Nevada-Cascades, westward  M S L I H

Epargyreus clarus Rocky Mountain segregate ,  NE OR, E WA, ID, MT, WY, C & N UT, CO, N NM, Black Hills  M S

Epargyreus clarus clarus (Cramer, 1775)  E USA and Canada to far NE Mexico (hypothetical)  M S HS L1 L2 L3 L4 I H G

Epargyreus clarus huachuca Dixon, 1955  S AZ, S NM, far NW Mexico  M T S L I H

Epargyreus clarus profugus Austin, 1998  S NV, N AZ, possibly to central AZ-NM  M T L

Epargyreus antaeus (Hewitson, 1867)  Jamaican Silverdrop  Jamaica  M T S L G

Epargyreus spanna Evans, 1952  Hispaniolan Silverdrop  Hispaniola  M T S G

Epargyreus barisses (Hewitson, 1874)  TL: Bolivia

Epargyreus barisses barisses (Hewitson, 1874)  TL: Bolivia  M T S G

Epargyreus barisses argentina Mabille, 1903  TL: ?  M T S G

Epargyreus tmolis (Burmeister, 1875)  TL: Argentina  M S G

Epargyreus socus (Hübner, [1825])  TL: ?

Epargyreus socus orizaba Scudder, 1872  E Mexico to Guatemala  M T S L G

Epargyreus socus chota Evans, 1952  TL: Trinidad  M T

Epargyreus socus dicta Evans, 1952  TL: Bolivia  M T

Epargyreus socus sinus Evans, 1952  TL: Brazil (Pa)  M T S

Epargyreus socus socus (Hübner, [1825])  TL: ?  M G

Epargyreus socus pseudexadeus Westwood, 1852  TL: Brazil ([MG])  M T S

Epargyreus windi H. Freeman, 1969  Wind’s Silverdrop  W Mexico  M T S L G

Epargyreus exadeus (Cramer, 1779)  TL: Surinam

Epargyreus exadeus cruza Evans, 1952  (S TX? [CA?]as stray), E & W Mexico to Panama  M T IL L G

Epargyreus exadeus exadeus (Cramer, 1779)  TL: Surinam  M S G

Epargyreus nutra Evans, 1952  TL: Colombia  M T G

Epargyreus aspina Evans, 1952  Spineless Silverdrop  E & W Mexico to Venezuela and Ecuador  M T L G

Epargyreus spina Evans, 1952  Spined Silverdrop  TL: Colombia

Epargyreus spina spina Evans, 1952  E & W Mexico to Brazil and Bolivia  M T S G

Epargyreus spina verruga Evans, 1952  TL: Peru  M T

Epargyreus spinta Evans, 1952  TL: Colombia  M T S

Epargyreus spinosa Evans, 1952  Suffused Silverdrop  W Mexico to Panama  M T S G

Epargyreus clavicornis (Herrich-Schäffer, 1869)  TL: ?

Epargyreus clavicornis gaumeri Godman & Salvin, 1893  SE Mexico (?) to Honduras  M T OI G

Epargyreus clavicornis tenda Evans, 1955  SE Mexico to Venezuela and Ecuador  M T S L G

Epargyreus clavicornis clavicornis (Herrich-Schäffer, 1869)  TL: ?  M T IL G

Epargyreus brodkorbi H. Freeman, 1969  Barely-spotted Silverdrop  Chiapas  M T S L G

Epargyreus deleoni H. Freeman, 1977  Long-spotted Silverdrop  Yucatan Peninsula (CAMP, QROO, YUC Mexico) and Belize  M T S G

Epargyreus enispe (Hewitson, 1867)  TL: S. Amer.

Epargyreus enispe enispe (Hewitson, 1867)  TL: S. Amer.  M T S G

Epargyreus enispe elta Evans, 1952  TL: Bolivia  M T

Polygonus  Hübner, [1825]

Polygonus leo (Gmelin, [1790])  Hammock Skipper  TL: America

Polygonus leo arizonensis (Skinner, 1911)  SW USA (mostly as strays) & S TX to Central America  M T S L I1 I2 G

Polygonus leo pallida Röber, 1925  Central America to Argentina  M T S L

Polygonus leo histrio Röber, 1925  Florida and Cuba, some Bahamian islands  M S L I

Polygonus leo leo (Gmelin, [1790])  [= lividus, = ishmael], Hispaniola, Puerto Rico, SE to St. Lucia  M T S IL L G

Polygonus leo hagar Evans, 1952  Jamaica  M T S

Polygonus savigny (Latreille, [1824])  Manuel’s Skipper  TL: Antilles

Polygonus savigny savigny (Latreille, [1824])  [= manueli], continental, (S TX as stray), N Mexico to Argentina  M T S1 S2 HS L I G

Polygonus savigny punctus E. Bell & W. Comstock, 1948  Lesser Antilles; St. Martin S to Grenadines  M T S

Chioides  Lindsey, 1921

Chioides albofasciatus (Hewitson, 1867)  White-striped Longtail  S TX, S NM, SE AZ, to Costa Rica; Baja California Sur  M T S L1 L2 L3 I H G

Chioides catillus (Cramer, 1779)  TL: Surinam

Chioides catillus albius Evans, 1952  Costa Rica and Panama  M T S1 S2 L I G

Chioides catillus catillus (Cramer, 1779)  TL: Surinam  M S1 S2 L

Chioides catillus jethira (A. Butler, 1870)  TL: Peru  M T S G

Chioides churchi E. Bell & W. Comstock, 1948  Jamaican Longtail  Jamaica  M T S

Chioides cinereus (Mabille & Vuillot, 1891)  TL: Brazil  M T S IL

Chioides concinnus (Mabille, 1877)  TL: Brazil  M T OI S G

Chioides vintra Evans, 1952  St. Vincent Longtail  St. Vincent, Grenadines, Grenada  M T S

Chioides iverna Evans, 1952  TL: Peru  M T S

Chioides zilpa (A. Butler, 1872)  Zilpa Longtail  S AZ-NM (stray) & S TX, S to Venezuela and Ecuador  M T OD S1 S2 L I1 I2 G

Chioides ixion (Plötz, 1880)  Hispaniolan Longtail  Hispaniola  M T S

Chioides marmorosa (Herrich-Schäffer, 1865)  Cuban Longtail  Cuba  M T S

Aguna  R. Williams, 1927

Aguna asander (Hewitson, 1867)  Gold-spotted Aguna  TL: Brazil (Am)

Aguna asander asander (Hewitson, 1867)  SE AZ & S TX (as stray) to Argentina  M T S L I G

Aguna asander haitensis (Mabille & Boullet, 1912)  Cuba and Hispaniola  M T S

Aguna asander jasper Evans, 1952  Jamaica  M T S L

Aguna megaeles (Mabille, 1888)  TL: [Brazil (SC)]

Aguna megaeles malia Evans, 1952  TL: Venezuela  M T G

Aguna megaeles megaeles (Mabille, 1888)  TL: [Brazil (SC)]  M T OI S IL

Aguna albistria (Plötz, 1880)  TL: Brazil ([Pa])

Aguna albistria albistria (Plötz, 1880)  TL: Brazil ([Pa])  M T TI S L

Aguna albistria leucogramma (Mabille, 1888)  SE Mexico to Colombia and Venezuela (Surinam?)  M T OI S

Aguna metophis (Latreille, [1824])  Long-tailed Aguna  S Texas to S Brazil  M T S L I

Aguna camagura (R. Williams, 1926)  TL: Brazil (MT)  M T S

Aguna claxon Evans, 1952  Emerald Aguna  Cuba, S Texas to Colombia and Surinam  M T S L I G

Aguna clina Evans, 1952  TL: Colombia  M T S

Aguna latifascia Austin & O. Mielke, 1998  TL: Ecuador  M S

Aguna coelus (Stoll, 1781)  Stoll's Aguna  Costa Rica and Panama to Peru and central Brazil  M T S

Aguna cirrus Evans, 1952  TL: Brazil (SP)  M T S G

Aguna coeloides Austin & O. Mielke, 1998  Austin's Aguna  SE Mexico to southern Brazil  M T S1 S2 I1 I2

Aguna nicolayi Austin & O. Mielke, 1998  TL: Brazil (MT)  M T S

Aguna venezuelae O. Mielke, 1971  TL: Venezuela  M S

Aguna latimacula Austin & O. Mielke, 1998  TL: Brazil (MG)  M T

Aguna aurunce (Hewitson, 1867)  TL: Brazil (Am)

Aguna aurunce aurunce (Hewitson, 1867)  TL: Brazil (Am)  M T S

Aguna aurunce hypozonius (Plötz, 1880)  SE Mexico to Venezuela  M S L I

Aguna glaphyrus (Mabille, 1888)  TL: Brazil (SC)  M T S

Aguna penicillata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna spicata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna longicauda Austin & O. Mielke, 1998  TL: Brazil (Rd)  M OI

Aguna ganna (Möschler, 1879)  Ganna Aguna  Costa Rica to Colombia and Venezuela  M T S IL

Aguna panama Austin & O. Mielke, 1998  Panamanian Aguna  Honduras to Venezuela  M T1 T2 S I

Aguna spatulata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna similis Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna mesodentata Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna squamalba Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Aguna parva Austin & O. Mielke, 1998  TL: Brazil (Rd)  M T S

Typhedanus  A. Butler, 1870

Typhedanus crameri McHenry, 1960  TL: not specified  M T S L

Typhedanus undulatus (Hewitson, 1867)  Mottled Longtail  S Texas (as stray) to Argentina  M T S1 S2 HS L I G

Typhedanus galbula (Plötz, 1880)  TL: Brazil ([Pa])  M T

Typhedanus optica Evans, 1952  TL: Guyana

Typhedanus optica optica Evans, 1952  TL: Guyana  M T S

Typhedanus optica goya Evans, 1952  TL: Brazil (Go)  M T S

Typhedanus stylites (Herrich-Schäffer, 1869)  TL: [Brazil]  M T

Typhedanus aziris (Hewitson, 1867)  TL: Brazil (RJ)  M T

Typhedanus umber (Herrich-Schäffer, 1869)  TL: ?  M T

Typhedanus ampyx (Godman & Salvin, 1893)  Gold-tufted Skipper  W & NE Mexico to Costa Rica  M T S1 S2 L I G

Typhedanus salas H. Freeman, 1977  Salas Skipper  S Mexico & Belize  M T L

Typhedanus cursinoi O. Mielke, 1979  TL: Brazil (Ba)  M OI G

Typhedanus eliasi O. Mielke, 1979  TL: Brazil (ES)  M OI G

Polythrix  E. Watson, 1893

Polythrix octomaculata (Sepp, [1844])  Eight-spotted Longtail  (SE AZ as stray), S Texas to Argentina  M T OI S1 S2 L I G

Polythrix maizae Hellebuyck, 1998  Melanic Longtail  Nicaragua  M T

Polythrix mexicanus H. Freeman, 1969  Mexican Longtail  SE AZ, S TX (as stray), N Mexico to Costa Rica  M T S1 S2 L

Polythrix roma Evans, 1952  TL: Brazil (Pa)  M T

Polythrix asine (Hewitson, 1867)  Asine Longtail  N Mexico to Panama  M T S1 S2 IL L I1 I2

Polythrix hirtius (A. Butler, 1870)  TL: Venezuela  M T

Polythrix gyges Evans, 1952  TL: Peru  M T S

Polythrix minvanes (R. Williams, 1926)  TL: Brazil (MT)  M T G

Polythrix ceculus (Herrich-Schäffer, 1869)  TL: ?  M T

Polythrix caunus (Herrich-Schäffer, 1869)  Four-spotted Longtail  E Mexico to S Brazil and Paraguay  M T S1 S2 L I G

Polythrix auginus (Hewitson, 1867)  Auginus Longtail  SE Mexico to S Brazil  M T S IL L I

Polythrix metallescens (Mabille, 1888)  Metallescens Skipper  SE Mexico to Peru and northern Brazil  M T S L G

Polythrix kanshul Shuey, 1991  Kanshul Longtail  S Mexico, Belize, NE Costa Rica  M S L I1 I2

Polythrix eudoxus (Stoll, 1781)  TL: Surinam  M OI L

Cephise  Evans, 1952

Cephise cephise (Herrich-Schäffer, 1869)  TL: ?  M T S

Cephise impunctus Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise glarus (Mabille, 1888)  TL: Brazil (Pa)  M T OI S

Cephise nuspesez Burns, 1996  Burns' Scarlet-eye  E Mexico to Costa Rica  M T S1 S2 I1 I2

Cephise malesedis Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise maculatus Austin & O. Mielke, 2000  TL: Brazil (Rd)  M T

Cephise burnsi Austin & O. Mielke, 2000  TL: Brazil (ES)  M OI

Cephise mexicanus Austin & O. Mielke, 2000  Mexican Scarlet-eye  SLP, OAX (Mexico)  M T

Cephise procerus (Plötz, 1880)  TL: Brazil (Pa)  M T

Cephise aelius (Plötz, 1880)  Longtailed Scarlet-eye  E & S Mexico to Venezuela and Colombia  M T S1 S2 L I1 I2 G

Cephise callias (Mabille, 1888)  TL: [Venezuela]  M T OI S IL

Cephise guatemalaensis (H. Freeman, 1977)  Guatemalan Scarlet-eye  Guatemala, Chiapas, Quintana Roo  M T

Venada  Evans, 1952

Venada [n. sp.] N/A, N/A  Cryptic Scarlet-eye  ["advena"], Mexico, Guatemala, (Nicaragua?)  M S

Venada nevada (Burns, 2005)  Nevada Scarlet-eye  Costa Rica  M OD S1 S2 I1 I2 G

Venada advena (Mabille, 1889)  Advena Scarlet-eye  Panama and Colombia, at least  M T OI

Venada daneva (Burns, 2005)  Daneva Scarlet-eye  Costa Rica  M OD S1 S2 L I1 I2 G

Venada cacao (Burns, 2005)  Cacao Scarlet-eye  Costa Rica and Panama  M T OD S I G

Venada naranja (Burns, 2005)  Orange Scarlet-eye  Costa Rica  M T OD S1 S2 I1 I2 G

Heronia  Mabille & Boullet, 1912

Heronia labriaris (A. Butler, 1877)  TL: Brazil (Am)  M T S IL

Chrysoplectrum  E. Watson, 1893

Chrysoplectrum pervivax (Hübner, [1819])  Pervivax Skipper  Costa Rica to Bolivia, Peru and N Brazil  M T OI S1 S2 HS I

Chrysoplectrum bahiana (Herrich-Schäffer, 1869)  TL: [Brazil (Ba)]  M T OI S

Chrysoplectrum orphne (Plötz, 1881)  TL: Brazil (RJ)  M T TI

Chrysoplectrum otriades (Hewitson, 1867)  TL: Brazil (Am)  M T S

Chrysoplectrum epicincea (A. Butler & H. Druce, 1872)  Stub-tailed Skipper  SE Mexico to Costa Rica  M T S G

Chrysoplectrum perniciosus (Herrich-Schäffer, 1969)  Perniciosus Skipper  Panama to Bolivia and C Brazil  M T S L

Chrysoplectrum cuminaensis R.F. d'Almeida, 1976  TL: Brazil (Pa)  M S

Chrysoplectrum orpheus (Plötz, 1881)  TL: Brazil (Pa)  M T TI

Chrysoplectrum perna Evans, 1952  TL: Colombia  M T

Chrysoplectrum albovenae E. Bell, 1932  TL: Brazil (SC)  M T

Zestusa  Lindsey, 1925

Zestusa dorus (W. H. Edwards, 1882)  Short-tailed Skipper (Northern Zestusa)  far S CO-UT, AZ, NM, W TX, N Mexico  M T S L1 L2 I Ha G

Zestusa elwesi (Godman & Salvin, 1893)  Mexican Zestusa  Durango to Oaxaca  M T S L G

Zestusa staudingeri (Mabille, 1888)  Southern Zestusa  Chiapas to El Salvador  M T OI S L

Zestusa levona Steinhauser, 1972  Levona's Zestusa  Chiapas to El Salvador  M T

Codatractus  Lindsey, 1921

Codatractus carlos Evans, 1952  Carlos' Mottled-Skipper  TL: Honduras

Codatractus carlos carlos Evans, 1952  C Mexico to Colombia (at least)  M T S L G

Codatractus carlos arguta Evans, 1952  TL: Trinidad  M T

Codatractus carlos rowena Evans, 1952  TL: Venezuela  M T

Codatractus alcaeus (Hewitson, 1867)  White-crescent Mottled-Skipper  TL: Nicaragua

Codatractus alcaeus alcaeus (Hewitson, 1867)  S Texas to Costa Rica  M T S L I G

Codatractus alcaeus apulia Evans, 1952  Panama (?) to W Ecuador  M T S G

Codatractus aminias (Hewitson, 1867)  TL: Brazil (MG)  M T L

Codatractus yucatanus H. Freeman, 1977  Yucatan Mottled-Skipper  Yucatan Peninsula  M T

Codatractus arizonensis (Skinner, 1905)  Arizona Mottled-Skipper  S AZ, S NM, W TX to Oaxaca  M T S L1 L2 I H G

Codatractus sallyae A. Warren, 1995  Sally’s Mottled-Skipper  W Mexico, Yucatan Peninsula, Guatemala  M T S L

Codatractus melon (Godman & Salvin, 1893)  Melon Mottled-Skipper  N Mexico to Costa Rica  M T S1 S2 L I

Codatractus bryaxis (Hewitson, 1867)  Golden Mottled-Skipper  N Mexico to Honduras  M T S L G

Codatractus imalena (A. Butler, 1872)  Bluish Mottled-Skipper  Costa Rica to northern Brazil  M T S1 S2 L I1 I2

Codatractus cyledis (Dyar, 1912)  Cyledis Mottled-Skipper  S Mexico  M T L Ha G

Codatractus cyda (Godman, 1901)  Cyda Mottled-Skipper  Chiapas to Nicaragua  M T OI

Codatractus uvydixa (Dyar, 1914)  Variable Mottled-Skipper  S Mexico: Guerrero to Chiapas

Codatractus uvydixa dark segregate ,  W Mexico: Sonora to Michoacan  M S L I

Codatractus uvydixa uvydixa (Dyar, 1914)  TL: Mexico (Gue)  M T S L Ha

Codatractus valeriana (Plötz, 1881)  Valeriana Skipper  [= mysie], SE Arizona to Oaxaca  M T S HS L I H F Ha G

Codatractus hyster (Dyar, 1916)  Hyster Skipper  Mexico, El Salvador  M T S L G

Ridens  Evans, 1952

Ridens crison (Godman & Salvin, 1893)  Many-spotted Ridens  TL: Guatemala

Ridens crison crison (Godman & Salvin, 1893)  SE Mexico, Guatemala  M T S G

Ridens crison howarthi Steinhauser, 1974  El Salvador  M T G

Ridens crison cachinnans (Godman, 1901)  Costa Rica, Panama  M T OI S I G

Ridens ridens (Hewitson, 1876)  Frosted Skipper  Costa Rica & Panama  M T S G

Ridens fieldi Steinhauser, 1974  Field's Ridens  Guatemala (Volcan Sta. Maria)  M T G

Ridens toddi Steinhauser, 1974  Todd’s Skipper  S Mexico (CHIS) to El Salvador  M T S G

Ridens biolleyi (Mabille, 1900)  Biolley's Ridens  Costa Rica, Panama  M T S1 S2 I1 I2 I3

Ridens mercedes Steinhauser, 1983  White-tailed Ridens  SW Mexico  M T S G

Ridens harpagus (C. Felder & R. Felder, 1867)  TL: Colombia  M T S

Ridens philistus (Hopffer, 1874)  TL: Peru

Ridens philistus philistus (Hopffer, 1874)  TL: Peru  M T

Ridens philistus philia Evans, 1952  TL: Colombia  M T

Ridens mephitis (Hewitson, 1876)  Hewitson's Ridens  SE Mexico to Peru and Bolivia  M T S1 S2 I1 I2

Ridens fulminans (Herrich-Schäffer, 1869)  TL: ?  M T S

Ridens fulima Evans, 1952  TL: Brazil (ES)  M T S L

Ridens panche (R. Williams, 1927)  Panche Ridens  Costa Rica to Colombia and Ecuador  M T S1 S2 I1 I2

Ridens pacasa (R. Williams, 1927)  TL: Bolivia  M T S

Ridens bidens Austin, 1998  TL: Brazil (Rd)  M OI

Ridens allyni H. Freeman, 1979  Allyn’s Ridens  SE Mexico, Guatemala  M T1 T2 G

Ridens miltas (Godman & Salvin, 1893)  Mexican Ridens  E Mexico  M T S G

Ridens bridgmani (Weeks, 1902)  TL: Bolivia  M T S

Ridens nora Evans, 1952  TL: Peru  M T S

Ridens tristis (Draudt, [1922])  TL: [Bolivia]  M T S G

Ridens telegonoides (Mabille & Boullet, 1912)  TL: Colombia  M T OI

Urbanus  Hübner, [1807]

Urbanus proteus (Linnaeus, 1758)  Long-tailed Skipper  TL: America

Urbanus proteus proteus (Linnaeus, 1758)  continental; E USA, S CA, S AZ, S NM, TX to Argentina  M T S1 S2 HS IL L I1 I2 I3 H G

Urbanus proteus domingo (Scudder, 1872)  Bahamas and Antilles  M S

Urbanus viterboana (Ehrmann, 1907)  Bluish Longtail  E & W Mexico to Colombia and Ecuador  M T S1 S2 L I G

Urbanus dubius Steinhauser, 1981  TL: Colombia  M T

Urbanus belli (Hayward, 1935)  Bell's Longtail  W Mexico & S Texas (as stray) to Argentina  M T TI S1 S2 L I1 I2 G

Urbanus huancavillcas (R. Williams, 1926)  TL: Ecuador  M T S

Urbanus pronta Evans, 1952  Spot-banded Longtail  E & W Mexico to Argentina  M T S1 S2 L I1 I2

Urbanus longicaudus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus parvus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus pronus Evans, 1952  Pronus Longtail  (S TX one-time stray), E & S Mexico to Argentina  M T S L

Urbanus esmeraldus (A. Butler, 1877)  Esmeralda Longtail  (S TX as stray), E & W Mexico to Peru & Guianas, Trinidad  M T S1 S2 L I G

Urbanus esma Evans, 1952  Esma Longtail  Panama to Brazil, Trinidad  M T S

Urbanus evona Evans, 1952  Turquoise Longtail  E & W Mexico to Colombia  M T S1 S2 L I G

Urbanus elmina Evans, 1952  Andean Longtail  (Costa Rica?), Colombia, Ecuador  M T S

Urbanus viridis H. Freeman, 1970  Rare Longtail  SE Mexico (VER)  M T

Urbanus esta Evans, 1952  Esta Longtail  E & W Mexico to Ecuador and Brazil  M T S1 S2 L I

Urbanus prodicus E. Bell, 1956  Montane Longtail  E & W Mexico to Costa Rica; often in cloud forest  M T S L

Urbanus magnus Steinhauser, 1981  TL: Ecuador  M T

Urbanus velinus (Plötz, 1880)  Velinus Skipper  (SE Mexico?). Guatemala to Brazil and Trinidad  M T S

Urbanus villus Austin, 1998  TL: Brazil (Rd)  M OI

Urbanus dorantes (Stoll, 1790)  Dorantes Longtail  TL: Surinam

Urbanus dorantes calafia (R. Williams, 1926)  Baja California Sur  M T S L

Urbanus dorantes dorantes (Stoll, 1790)  continental; Florida, S AZ, S NM, TX to Argentina  M T S HS L1 L2 L3 I1 I2 H G

Urbanus dorantes cramptoni W. Comstock, 1944  Hispaniola, Puerto Rico, Mona, Culebra, Virgin Is., Gr. Cayman  M T S

Urbanus dorantes santiago (Lucas, 1857)  Cuba, N Andros, Mangrove Cay, Grand Bahama  M T S

Urbanus dorantes galapagensis (F. Williams, 1911)  TL: Galapagos  M T

Urbanus obscurus (Hewitson, 1867)  Dark Longtail  Lesser Antilles; St. Martin S to Grenadines, Barbados  M T S L

Urbanus teleus (Hübner, 1821)  Teleus Longtail  S Texas to Argentina  M T S1 S2 Ab IL L1 L2 I G

Urbanus tanna Evans, 1952  Tanna Longtail  S Texas (as stray) to Ecuador, Guianas, Trinidad  M T S L G

Urbanus ambiguus de Jong, 1983  TL: Surinam  M T

Urbanus cindra Evans, 1952  TL: Bolivia  M T S

Urbanus zagorus (Plötz, 1880)  TL: Brazil (RS)  M T

Urbanus simplicius (Stoll, 1790)  Plain Longtail  S Texas (as stray) to Argentina  M T S L1 L2 I1 I2 G

Urbanus procne (Plötz, 1881)  Brown Longtail  S Texas, SE AZ/SW NM (as stray), to Argentina  M S L1 L2 I G

Urbanus evenus (Ménétriés, 1855)  TL: Brazil  M S IL

Urbanus trebia (Möschler, 1879)  White-capped Longtail  El Salvador to Colombia and Trinidad  M T S G

Urbanus carmelita (Herrich-Schäffer, 1869)  Carmelita Longtail  TL: [Brazil]

Urbanus carmelita carmelita (Herrich-Schäffer, 1869)  Panama to Brazil  M T S IL

Urbanus carmelita barra Evans, 1952  TL: Brazil (Ba)  M OD

Urbanus reductus (N. Riley, 1919)  TL: Brazil (Am)  M T S

Urbanus doryssus (Swainson, 1831)  White-tailed Longtail  TL: Brazil (Ba)

Urbanus doryssus chales (Godman & Salvin, 1893)  W Mexico  M T S L G

Urbanus doryssus E Mexican segregate ,  ["d. doryssus"], S Texas to El Salvador  M S L1 L2

Urbanus doryssus doryssus (Swainson, 1831)  Chiapas to S Brazil & Bolivia  M T OI OD S L I1 I2

Urbanus doryssus albicuspis (Herrich-Schäffer, 1869)  TL: ?  M T

Urbanus albimargo (Mabille, 1876)  White-edged Longtail  TL: Panama; Colombia?

Urbanus albimargo albimargo (Mabille, 1876)  NE Mexico to Venezuela and Ecuador  M T S1 S2 IL L I

Urbanus albimargo takuta Evans, 1952  TL: Guyana  M T S

Urbanus albimargo rica Evans, 1952  TL: Paraguay  M T

Urbanus virescens (Mabille, 1877)  Green-washed Longtail  Panama to S Brazil and Paraguay  M T S L G

Urbanus chalco (Hübner, 1823)  Great White Longtail  Costa Rica to S Brazil and Paraguay  M S IL L G

Astraptes  Hübner, [1819]

Astraptes talus (Cramer, 1777)  Green Flasher  E & S Mexico to Argentina; Jamaica, Cuba, Hispaniola, etc.  M T S L G

Astraptes fulgerator (Walch, 1775)  TL: ?

Astraptes fulgerator azul (Reakirt, [1867])  This appears to be a complex of continental species-level taxa.  M T S L1 L2 I H

Astraptes fulgerator fulgerator (Walch, 1775)  TL: ?  M OI G

Astraptes fulgor (Hayward, 1939)  TL: Argentina  M TI

Astraptes naxos (Hewitson, 1867)  TL: Brazil  M T S

Astraptes tucuti (R. Williams, 1927)  Tucuti Flasher  SE Mexico to Peru and Bolivia  M T OI S L

Astraptes brevicauda (Plötz, 1886)  Short-tailed Flasher  [= talthybius], SE Mexico to Costa Rica  M T S1 S2 L I

Astraptes palliolum (H. Druce, 1908)  Ochre-stained Flasher  Costa Rica  M T OI S

Astraptes egregius (A. Butler, 1870)  Small-spotted Flasher  TL: ?

Astraptes egregius egregius (A. Butler, 1870)  S TX (rare stray) to Colombia and Venezuela  M T S L Ha G

Astraptes egregius coxeyi (R. Williams, 1931)  TL: Ecuador  M T

Astraptes erycina (Plötz, 1881)  TL: Brazil  M S IL G

Astraptes halesius (Hewitson, 1877)  TL: French Guiana  M T S

Astraptes mabillei Steinhauser, 1989  TL: Bolivia  M S

Astraptes phalaecus (Godman & Salvin, 1893)  Yellow-edged Flasher  E & W Mexico to Colombia  M T S L G

Astraptes apastus (Cramer, 1777)  Broad-banded Flasher  TL: Surinam

Astraptes apastus apastus (Cramer, 1777)  SE Mexico to Guianas and N Brazil  M S L I G

Astraptes apastus pusa Evans, 1952  TL: Brazil (Ba)  M T

Astraptes aulus (Plötz, 1881)  Green-tinged Flasher  Nicaragua to Peru and Argentina  M T OI S G

Astraptes enotrus (Stoll, 1781)  White-spotted Flasher  E & W Mexico to S Brazil, Paraguay, Bolivia  M T S L G

Astraptes janeira (Schaus, 1902)  Schaus' Flasher  TL: Brazil (RJ)  M T S L

Astraptes xagua (Lucas, 1857)  Antillean Flasher  TL: Cuba

Astraptes xagua xagua (Lucas, 1857)  Cuba, Andros  M T S

Astraptes xagua christyi (Sharpe, 1898)  Hispaniola  M T S

Astraptes megalurus (Mabille, 1877)  Long-tailed Flasher  E & W Mexico to Colombia  M T S L G

Astraptes elorus (Hewitson, 1867)  TL: ?  M T S L

Astraptes alardus (Stoll, 1790)  TL: Surinam

Astraptes alardus latia Evans, 1952  S TX (rare stray), E Mexico to Colombia  M T S1 S2 L I Ha

Astraptes alardus aquila Evans, 1952  TL: Colombia  M T

Astraptes alardus alardus (Stoll, 1790)  TL: Surinam  M T OD S L G

Astraptes habana (Lucas, 1857)  Frosty Flasher  TL: Cuba

Astraptes habana habana (Lucas, 1857)  Cuba, Isla de Juventud  M S

Astraptes habana heriul (Mabille & Boullet, 1912)  Hispaniola  M T S

Astraptes alector (C. Felder & R. Felder, 1867)  TL: Colombia

Astraptes alector hopfferi (Plötz, 1881)  [= gilberti], S TX (stray), E & W Mexico to NW Brazil and Bolivia  M T S L

Astraptes alector alector (C. Felder & R. Felder, 1867)  TL: Colombia  M T

Astraptes cretatus (Hayward, 1939)  Cretatus Flasher  TL: Ecuador

Astraptes cretatus cretatus (Hayward, 1939)  [= alfius], Nicaragua, Colombia to Bolivia  M T G

Astraptes cretatus adoba Evans, 1952  TL: Brazil (ES)  M T S

Astraptes creteus (Cramer, 1780)  Whitened Flasher  TL: Surinam

Astraptes creteus crana Evans, 1952  SE Mexico to Panama  M T S

Astraptes creteus crilla Evans, 1952  TL: Ecuador  M T

Astraptes creteus cyprus Evans, 1952  TL: Bolivia  M T

Astraptes creteus creteus (Cramer, 1780)  Panama to Guianas and Bolivia  M S IL G

Astraptes creteus siges (Mabille, 1903)  TL: Brazil  M T

Astraptes bifascia (Herrich-Schäffer, 1869)  Green-headed Flasher  SE Mexico to Panama  M T S

Astraptes latimargo (Herrich-Schäffer, 1869)  White-edged Flasher  Panama, Colombia, Ecuador  M T OI S

Astraptes tinda Evans, 1952  TL: Brazil (Pa)  M T

Astraptes chiriquensis (Staudinger, 1876)  Chiriqui Flasher  TL: Panama

Astraptes chiriquensis chiriquensis (Staudinger, 1876)  E Mexico to Venezuela and Ecuador  M T S IL L

Astraptes chiriquensis erana Evans, 1952  TL: Ecuador  M T

Astraptes chiriquensis meretrix (Hewitson, 1876)  TL: Ecuador  M T S

Astraptes chiriquensis oenander (Hewitson, 1876)  TL: Brazil (Pa)  M IL G

Astraptes weymeri (Plötz, 1882)  Weymer’s Flasher  TL: ?  M S HS

Astraptes jaira (A. Butler, 1870)  Jamaican Flasher  Jamaica  M T S L

Astraptes cassander (Fabricius, 1793)  Cuban Flasher  Cuba and Isla de Juventud  M T S G

Astraptes galesus (Mabille, 1888)  TL: Peru

Astraptes galesus cassius Evans, 1952  Costa Rica to Venezuela  M T S L G

Astraptes galesus galesus (Mabille, 1888)  TL: Peru  M T OI

Astraptes anaphus (Cramer, 1777)  TL: Surinam

Astraptes anaphus annetta Evans, 1952  S Texas to Guiana and Peru  M T S1 S2 L1 L2 I G

Astraptes anaphus anaphus (Cramer, 1777)  TL: Surinam  M T

Astraptes anaphus anoma Evans, 1952  TL: Trinidad  M T

Astraptes anaphus aniza Evans, 1952  TL: Peru  M T

Astraptes anaphus anausis (Godman & Salvin, 1896)  Greater Antilles, Virgin Is., Montserrat to Grenada  M T S

Narcosius  Steinhauser, 1986

Narcosius narcosius (Stoll, 1790)  TL: Surinam  M T S G

Narcosius aulina (Evans, 1952)  TL: French Guiana  M T

Narcosius samson (Evans, 1952)  Samson Flasher  SE Mexico to Brazil and Bolivia  M T S1 S2 I1 I2 G

Narcosius mura (R. Williams, 1927)  TL: Brazil (Am)  M T S IL G

Narcosius pseudomura Austin, 1996  TL: Brazil (Rd)  M OI G

Narcosius colossus (Herrich-Schäffer, 1869)  Colossal Flasher  TL: Venezuela

Narcosius colossus colossus (Herrich-Schäffer, 1869)  SE Mexico to Colombia and Venezuela  M T S1 S2 I1 I2 G

Narcosius colossus granadensis (Möschler, 1879)  TL: Venezuela  M T S G

Narcosius dosula (Evans, 1952)  TL: Brazil (RS)  M T

Narcosius hercules (E. Bell, 1956)  TL: Bolivia  M T

Narcosius parisi (R. Williams, 1927)  TL: Brazil (MT)

Narcosius parisi helen (Evans, 1952)  E & W Mexico to Panama  M T S I G

Narcosius parisi parisi (R. Williams, 1927)  TL: Brazil (MT)  M T

Narcosius odysseus Austin, 1996  TL: Brazil (Rd)  M OI G

Narcosius nazaraeus Steinhauser, 1986  Nazareus Flasher  SE Mexico to Guianas, Brazil, Peru, Ecuador and Bolivia  M T S L G

Narcosius steinhauseri Austin, 1996  TL: Brazil (Rd)  M OI G

Calliades  Mabille & Boullet, 1912

Calliades oryx (C. Felder & R. Felder, 1862)  TL: Brazil (Am)  M T OI S

Calliades zeutus (Möschler, 1879)  Zeutus Banded-Skipper  SE Mexico to Guianas and Bolivia  M T S1 S2 L I G

Autochton  Hübner, 1823

Autochton cellus (Boisduval & Le Conte, [1837])  Golden Banded-Skipper  E-central & SE USA, SE AZ, SW NM, Mexico, S to El Salvador  M T OI OD S1 S2 S3 S4 IL L1 L2 I Ha G Txt

Autochton siermadror Burns, 1984  East-Mexican Banded-Skipper  NE Mexico; cloud forest  M T L G

Autochton pseudocellus (Coolidge & Clémence, [1910])  Sonoran Banded-Skipper  Mexico; apparently extirpated from S Arizona  M T S L Ha G

Autochton cincta (Plötz, 1882)  Chisos Banded-Skipper  W Texas to Guatemala  M T S L I Ha G

Autochton vectilucis (A. Butler, 1872)  Central American Banded-Skipper  Chiapas to Panama  M T OD S L G

Autochton integrifascia (Mabille, 1891)  TL: Brazil (SP/RS)  M S L

Autochton neis (Geyer, 1832)  Broad Banded-Skipper  E & W Mexico to S Brazil and Paraguay  M T S Ab L G

Autochton reflexus (Mabille & Boullet, 1912)  TL: Brazil (SC)  M T S

Autochton longipennis (Plötz, 1882)  Spike Banded-Skipper  E & S Mexico to S Brazil  M T S L I G

Autochton zarex (Hübner, 1818)  Sharp Banded-Skipper  E & S Mexico to Argentina  M T S1 S2 HS L I G

Autochton bipunctatus (Gmelin, [1790])  Gmelin's Banded-Skipper  SE Mexico to Bolivia and Brazil  M T S L I

Autochton sulfureolus (Mabille, 1883)  TL: Brazil  M G

Autochton itylus Hübner, 1823  TL: Surinam  M OI S L

Thessia  Steinhauser, 1989

Thessia athesis (Hewitson, 1867)  Athesis Cloudywing  Panama, Colombia, Venezuela to Tobago  M T S L G

Thessia jalapus (Plötz, 1881)  Jalapus Cloudywing  S Texas (rare stray) to Colombia  M T S1 S2 L I1 I2 G

Achalarus  Scudder, 1872

Achalarus toxeus (Plötz, 1882)  Coyote Cloudywing  W Mexico and S Texas to Panama  M T S1 S2 L I G

Achalarus albociliatus (Mabille, 1877)  White-edged Cloudywing  TL: Colombia; Panama; Guatemala

Achalarus albociliatus albociliatus (Mabille, 1877)  SE AZ & S TX (?) as stray, E & W Mexico to Costa Rica  M T S1 S2 L I G

Achalarus albociliatus leucophrys (Mabille, 1898)  Panama and Colombia  M T S

Achalarus albociliatus nocera (Plötz, 1882)  TL: Colombia  M T OI

Achalarus lyciades (Geyer, 1832)  Hoary Edge  E USA, (Veracruz)  M S1 S2 HS L1 L2 L3 I G

Achalarus casica (Herrich-Schäffer, 1869)  Desert Cloudywing  SW USA (SE AZ to C TX), Mexico  M T S L I H G

Achalarus tehuacana (Draudt, 1922)  Tehuacan Cloudywing  S TX (rare stray), E Mexico to Guatemala  M T OD S L G

Thorybes  Scudder, 1872

Thorybes confusis E. Bell, 1923  Confused Cloudywing  SE USA  M T1 T2 S1 S2 S3 L1 L2 L3 I G

Thorybes bathyllus (J. E. Smith, 1797)  Southern Cloudywing  E USA  M OI TI S1 S2 S3 HS L1 L2 L3 L4 G

Thorybes mexicana (Herrich-Schäffer, 1869)  Mexican Cloudywing  TL: [Mexico]

Thorybes mexicana aemilea (Skinner, 1893)  Cascade Mts. N CA to C OR  M T S L1 L2 Ha

Thorybes mexicana blanca Scott, 1981  White Mts., CA-NV  M S L1 L2

Thorybes mexicana nevada Scudder, 1872  S Rocky Mountains (S WY-ID to NM, AZ), CA-NV Sierra Nevada  M S L1 L2 Ha G

Thorybes mexicana dobra Evans, 1952  NW Mexico, SE Arizona, SW New Mexico  M T S L

Thorybes mexicana mexicana (Herrich-Schäffer, 1869)  C & W Mexico  M T S L Ha G

Thorybes mexicana ducia Evans, 1952  TL: Panama  M OD

Thorybes diversus E. Bell, 1927  Western Cloudywing  C & N Sierra Nevada and Siskiyou Mts., CA, far SW OR  M S L1 L2 L3 I G

Thorybes drusius (W. H. Edwards, [1884])  Drusius Cloudywing, White-fringed Cloudywing  SE AZ, SW NM, W TX, Mexico  M T S L I Ha

Thorybes pylades (Scudder, 1870)  Northern Cloudywing  TL: USA (Ma)

Thorybes pylades indistinctus Austin & J. Emmel, 1998  Baja California (N) to Washington, Cascades (& Ochocos) west  M T S L1 L2 Ha

Thorybes pylades BCS segregate ,  Baja California Sur  M S

Thorybes pylades S Rocky Mountain segregate ,  NE OR, S ID, MT, WY, W SD, W NE, N & C UT, CO, N NM, N AZ  M S

Thorybes pylades southwestern segregate ,  S AZ, S NM, W Mexico to Oaxaca  M S L1 L2 L3 L4 H

Thorybes pylades albosuffusa H. Freeman, 1943  W Texas, NE Mexico  M T S1 S2

Thorybes pylades pylades (Scudder, 1870)  Canada, E USA  M T S1 S2 L1 L2 L3 L4 I G

Cabares  Godman & Salvin, 1894

Cabares potrillo (Lucas, 1857)  Potrillo Skipper  TL: Cuba

Cabares potrillo potrillo (Lucas, 1857)  SE AZ (stray), S TX to Costa Rica; Jamaica, Cuba, Hispaniola  M T S1 S2 L I G

Cabares potrillo reducta Mabille & Boullet, 1919  Panama, Colombia, Venezuela, Trinidad  M S

Cabares rinta Evans, 1952  TL: Argentina  M T S

Spathilepia  A. Butler, 1870

Spathilepia clonius (Cramer, 1775)  Falcate Skipper  S TX (sporadic) to N Argentina  M T S1 S2 IL L I G

Oechydrus  E. Watson, 1893

Oechydrus chersis (Herrich-Schäffer, 1869)  TL: ?

Oechydrus chersis chersis (Herrich-Schäffer, 1869)  TL: ?  M T

Oechydrus chersis ochrilinea (Schaus, 1902)  TL: Peru  M T S

Oechydrus chersis rufus Evans, 1953  TL: Brazil (RJ)  M OD S

Oechydrus chersis evelinda (A. Butler, 1870)  TL: Brazil (RJ)  M T TI S G

Ectomis  Mabille, 1878

Ectomis cythna (Hewitson, 1878)  TL: ?

Ectomis cythna cythna (Hewitson, 1878)  TL: ?  M T L

Ectomis cythna ega Evans, 1953  TL: Brazil (Am)  M OD

Nerula  Mabille, 1888

Nerula fibrena (Hewitson, 1877)  TL: Brazil (Am)  M T OI

Nerula tuba Evans, 1953  TL: Brazil (Pa)  M OD

Marela  Mabille, 1903

Marela tamyroides (C. Felder & R. Felder, 1867)  TL: Brazil (Am)  M T S L

Marela tamyris Mabille, 1903  TL: Brazil (Am)

Marela tamyris tamyris Mabille, 1903  TL: Brazil (Am)  M T TI S L

Marela tamyris tamba Evans, 1953  TL: Peru  M S

Cogia  A. Butler, 1870

Cogia hassan A. Butler, 1870  TL: Brazil (Pa)

Cogia hassan hassan A. Butler, 1870  TL: Brazil (Pa)  M T

Cogia hassan evansi E. Bell, 1937  TL: Bolivia  M T

Cogia abdul Hayward, 1947  TL: Argentina  M TI G

Cogia cajeta (Herrich-Schäffer, 1869)  Yellow-haired Skipper  TL: ?

Cogia cajeta cajeta (Herrich-Schäffer, 1869)  SE Mexico to Nicaragua  M T S L G

Cogia cajeta eluina Godman & Salvin, 1894  W Mexico to Costa Rica  M T S1 S2 L I

Cogia troilus Mabille, 1898  TL: Bolivia  M T

Cogia punctilia Plötz, 1882  TL: Brazil ([Ba])  M T

Cogia cerradicola (O. Mielke, 1967)  TL: Brazil (Go)  M OI G

Cogia outis (Skinner, 1894)  Outis Skipper  NE Mexico (CHIH, COAH), Texas, S-C OK to far SW MO  M T S L I H Ha G

Cogia hippalus (W. H. Edwards, 1882)  Acacia Skipper  TL: USA (Az)

Cogia hippalus peninsularis L. Miller & MacNeill, 1969  Baja California Sur  M T S

Cogia hippalus hippalus (W. H. Edwards, 1882)  AZ, SW NM, W & S TX, W Mexico  M T S L I H G

Cogia hippalus hiska Evans, 1953  E Mexico to Costa Rica  M S1 S2 L I G

Cogia hippalus hester Evans, 1953  TL: Venezuela  M OD

Cogia azila Evans, 1953  TL: Bolivia  M OD

Cogia caicus (Herrich-Schäffer, 1869)  Gold-costa Skipper  TL: ?

Cogia caicus moschus (W. H. Edwards, 1882)  SE AZ. S NM to OAX  M S L I H G

Cogia caicus caicus (Herrich-Schäffer, 1869)  Chiapas to Costa Rica  M T S G

Cogia calchas (Herrich-Schäffer, 1869)  Mimosa Skipper  S Texas to N Argentina  M T S1 S2 L I G

Cogia grandis N. Riley, 1921  TL: Brazil (MT)  M T S

Cogia aventinus (Godman & Salvin, 1894)  Trimmed Skipper  W Mexico, Yucatan  M T S IL L1 L2 Ha

Cogia mala Evans, 1953  Guatemalan Skipper  S Mexico, Guatemala  M S HS

Cogia elaites (Hewitson, 1867)  TL: Brazil (MG)  M T

Paracogia  O. Mielke, 1977

Paracogia acanthopoda O. Mielke, 1977  TL: Brazil (Pr)  M OI G

Telemiades  Hübner, [1819]

Telemiades delalande (Latreille, [1824])  Delalande Skipper  Guatemala to Venezuela, Peru, upper Amazon  M T S L

Telemiades litanicus (Hewitson, 1876)  TL: Brazil (Am)  M T S

Telemiades vansa Evans, 1953  TL: Guyana  M OD

Telemiades centrites (Hewitson, 1870)  TL: Ecuador

Telemiades centrites contra Evans, 1953  TL: Ecuador  M OD

Telemiades centrites centrites (Hewitson, 1870)  TL: Ecuador  M S Txt

Telemiades gallius (Mabille, 1888)  Centrites Skipper  Costa Rica and Panama, mid-elevation rain forest  M T OI S I

Telemiades chrysorrhoea (Godman & Salvin, 1893)  Chrysorrhoea Skipper  Costa Rica and Panama, cloud forest  M T S1 S2 L I1 I2

Telemiades vespasius (Fabricius, 1793)  TL: Indiis  M T

Telemiades squanda Evans, 1953  TL: Brazil (RJ)  M S IL G

Telemiades trenda Evans, 1953  TL: Surinam  M OD

Telemiades nicomedes (Möschler, 1879)  Dark Telemiades  SE Mexico to Guianas and upper Amazon  M T S1 S2 I1 I2

Telemiades meris (Plötz, 1886)  Meris Telemiades  TL: Colombia

Telemiades meris meris (Plötz, 1886)  Panama & Colombia  M T S

Telemiades meris brazus E. Bell, 1949  TL: Brazil (SC)  M T TI

Telemiades choricus (Schaus, 1902)  Mexican Telemiades  W & S Mexico, Belize  M T S L G

Telemiades megallus Mabille, 1888  Orange Telemiades  SE Mexico (?) to Costa Rica  M T OI S1 S2 I1 I2 G

Telemiades sila Evans, 1953  Sila Telemiades  Nicaragua to Colombia, Venezuela and Ecuador  M S

Telemiades epicalus Hübner, [1819]  Hübner's Telemiades  Panama to S Brazil  M T S L G

Telemiades penidas (Hewitson, 1867)  Penidas Telemiades  Costa Rica to S Brazil  M T S IL L

Telemiades avitus (Stoll, 1781)  Yellow-spotted Telemiades  SE Mexico to N Amazon  M T OI S L I G

Telemiades antiope (Plötz, 1882)  Plötz's Telemiades  TL: Colombia; Brazil (Pa)

Telemiades antiope antiope (Plötz, 1882)  Nicaragua, Costa Rica, Colombia, S Brazil  M T S1 S2 I1 I2

Telemiades antiope tosca Evans, 1953  TL: Brazil (Am)  M OD L

Telemiades fides E. Bell, 1949  Small Telemiades  E & W Mexico to Colombia, Ecuador and upper Amazon  M T S1 S2 L I1 I2

Telemiades amphion (Geyer, 1832)  TL: "West Indies"

Telemiades amphion pekahia (Hewitson, 1868)  TL: Venezuela  M Txt

Telemiades amphion amphion (Geyer, 1832)  TL: "West Indies"  M T S

Telemiades amphion misitheus Mabille, 1888  TL: Peru  M T OI S L

Telemiades amphion marpesus (Hewitson, 1876)  TL: Brazil  M IL G

Telemiades laogonus (Hewitson, 1876)  TL: Brazil

Telemiades laogonus nicola (Plötz, 1882)  TL: Brazil (Pa)  M T

Telemiades laogonus laogonus (Hewitson, 1876)  TL: Brazil  M T

Telemiades corbulo (Stoll, 1781)  Corbulo Skipper  Guianas, Ecuador, Peru, Bolivia, upper and lower Amazon  M T S L G

Telemiades oiclus (Mabille, 1889)  Oiclus Skipper  [= cora], Honduras to Panama  M T S I1 I2

Bungalotis  E. Watson, 1893

Bungalotis erythus (Cramer, 1775)  Spotted Scarlet-eye  SE Mexico to Guianas, Brazil and Bolivia  M S1 S2 I1 I2 G

Bungalotis diophorus (Möschler, 1883)  Diophorus Scarlet-Eye  Costa Rica to Guianas and N Brazil  M T S1 S2 I1 I2

Bungalotis gagarini O. Mielke, 1967  Gagarin's Scarlet-Eye  TL: Brazil (Go)  M OI IL G

Bungalotis midas (Cramer, 1775)  White-cheeked Scarlet-eye  SE Mexico to S Brazil and Paraguay  M T OI S1 S2 IL L I1 I2 G

Bungalotis aureus Austin, 2008  Golden Scarlet-Eye  TL: Ecuador  M OI G

Bungalotis astylos (Cramer, 1780)  Dark-cheeked Scarlet-eye  SE Mexico to Guianas, Peru and S Brazil  M T S1 S2 L I1 I2 G

Bungalotis milleri H. Freeman, 1977  Miller's Scarlet-eye  SE Mexico to Costa Rica  M T OD G

Bungalotis quadratum (Sepp, [1845])  Pallid Scarlet-eye  TL: Surinam

Bungalotis quadratum quadratum (Sepp, [1845])  SE Mexico to Guianas  M T OI S1 S2 L I1 I2

Bungalotis quadratum barba Evans, 1952  TL: Brazil (Pa)  M T

Bungalotis sipa de Jong, 1983  Sipa Scarlet-eye  Surinam  M T G

Bungalotis clusia Evans, 1952  Clusia Scarlet-eye  TL: Brazil (Pa)  M T

Bungalotis borax Evans, 1952  Borax Scarlet-eye  TL: French Guiana  M T S G

Bungalotis lactos Evans, 1952  Lactos Scarlet-eye  TL: Peru  M T G

Salatis  Evans, 1952

Salatis canalis (Skinner, 1920)  Canal Scarlet-eye  Costa Rica to Ecuador  M T S1 S2 I1 I2

Salatis salatis (Stoll, 1782)  Variable Scarlet-eye  Nicaragua, Panama to Guianas, Peru and S Brazil  M T S L G

Salatis cebrenus (Cramer, 1777)  TL: Surinam  M T OI

Salatis fulvius (Plötz, 1882)  TL: Brazil (Pa)  M T S HS

Salatis scyrus (E. Bell, 1934)  TL: Peru  M T

Salatis flavomarginatus (Sepp, [1851])  TL: Surinam  M OI I

Sarmientoia  Berg, 1897

Sarmientoia dinka Evans, 1952  TL: ?  M T

Sarmientoia eriopis (Hewitson, 1867)  TL: Brazil (Pa)  M T S

Sarmientoia phaselis (Hewitson, 1867)  TL: Brazil  M T IL G

Sarmientoia faustinus (Burmeister, 1878)  TL: Argentina  M S IL G

Sarmientoia haywardi O. Mielke, 1967  TL: Bolivia  M OI IL G

Sarmientoia similis O. Mielke, 1967  TL: Brazil (Pa)  M OI IL G

Sarmientoia almeidai O. Mielke, 1967  TL: Brazil (MT)  M OI S G

Sarmientoia browni O. Mielke, 1967  TL: Brazil (DF)  M OI S G

Nicephellus  Austin, 2008

Nicephellus nicephorus (Hewitson, 1876)  Two-spotted Scarlet-eye  SE Mexico to Guianas, N & S Brazil, NE Peru  M T S IL I

Euriphellus  Austin, 2008

Euriphellus euribates (Stoll, 1782)  Euribates Scarlet-eye  TL: Surinam

Euriphellus euribates euribates (Stoll, 1782)  Costa Rica to Brazil and Bolivia  M T OI S G

Euriphellus euribates polygius (Latreille, [1824])  TL: ?  M T IL

Dyscophellus  Godman & Salvin, 1893

Dyscophellus phraxanor (Hewitson, 1876)  Big-spotted Scarlet-eye  TL: Colombia; Panama

Dyscophellus phraxanor lama Evans, 1952  SE Mexico to Honduras  M T S

Dyscophellus phraxanor phraxanor (Hewitson, 1876)  Nicaragua to Ecuador  M T S1 S2 L I G

Dyscophellus phraxanor mena Evans, 1952  TL: Ecuador  M T

Dyscophellus marian Evans, 1952  TL: Ecuador  M S

Dyscophellus porcius (C. Felder & R. Felder, 1862)  Fiery Scarlet-eye  TL: Brazil (Am)

Dyscophellus porcius porcius (C. Felder & R. Felder, 1862)  S Mexico to Paraguay and S Brazil  M T S L G

Dyscophellus porcius doriscus (Hewitson, 1867)  TL: Brazil ([RJ])  M T S

Dyscophellus sebaldus (Stoll, 1781)  Sebaldus Scarlet-eye  Costa Rica to Guianas and N Brazil  M OI S IL

Dyscophellus erythras (Mabille, 1888)  TL: Brazil (Pa)  M T OI S

Dyscophellus mielkei Austin, 2008  TL: Brazil (Rd)  M OI

Dyscophellus diaphorus (Mabille & Boullet, 1912)  Diaphorus Scarlet-eye  Costa Rica to French Guiana and N Brazil  M T OI S

Dyscophellus porsena (E. Bell, 1934)  TL: Peru  M T

Dyscophellus ramon Evans, 1952  Plain Scarlet-eye  SE Mexico to Panama  M T S1 S2 I1 I2 G

Dyscophellus ramusis (Stoll, 1781)  Subtle Scarlet-eye  Costa Rica to S Brazil  M S L

Dyscophellus damias (Plötz, 1882)  TL: Brazil  M T S

Nascus  E. Watson, 1893

Nascus phintias Schaus, 1913  Schaus’ Scarlet-eye  SE Mexico to French Guiana and Peru  M T S1 S2 I

Nascus phocus (Cramer, 1777)  Common Scarlet-eye  E & W Mexico to S Brazil, Paraguay and Argentina  M T S1 S2 L I1 I2 G

Nascus solon (Plötz, 1882)  TL: S. Amer.

Nascus solon corilla Evans, 1952  S Mexico to Colombia and Venezuela  M T S1 S2 I1 I2

Nascus solon solon (Plötz, 1882)  TL: S. Amer.  M S L

Nascus broteas (Cramer, 1780)  Broteas Scarlet-eye  E Mexico to Argentina, S Brazil and Paraguay  M T S1 S2 HS L I1 I2 G

Pseudonascus  Austin, 2008

Pseudonascus paulliniae (Sepp, [1842])  Least Scarlet-eye  S Mexico to Bolivia and S Brazil  M T OI S1 S2 I1 I2 G

Pseudonascus prax Evans, 1952  TL: Brazil (Pa)  M T

Porphyrogenes  E. Watson, 1893

Porphyrogenes boliva Evans, 1952  TL: Venezuela  M T

Porphyrogenes ferruginea (Plötz, 1883)  TL: Brazil (Ba)  M T S

Porphyrogenes despecta (A. Butler, 1870)  TL: Brazil (Pa)  M T S

Porphyrogenes omphale (A. Butler, 1871)  Blue Scarlet-eye  Costa Rica, Panama, Venezuela, Guianas to Bolivia  M T OI S1 S2 L G

Porphyrogenes pausias (Hewitson, 1867)  TL: Brazil (Am)  M T S

Porphyrogenes probus (Möschler, 1877)  TL: Surinam  M T

Porphyrogenes vulpecula (Plötz, 1882)  TL: [Brazil (Pa)]  M T S G

Porphyrogenes sororcula (Mabille & Boullet, 1912)  TL: French Guiana  M T OI S

Porphyrogenes specularis Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes spanda Evans, 1952  TL: Brazil (Pa)  M T S

Porphyrogenes stresa Evans, 1952  TL: Peru  M T

Porphyrogenes zohra (Möschler, 1879)  Zohra Scarlet-eye  Honduras, Nicaragua, Venezuela, Guianas, N Brazil (Para)  M T S G

Porphyrogenes sparus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes speciosus Austin & O. Mielke, 2008  Speciosus Scarlet-eye  Costa Rica  M T G

Porphyrogenes peterwegei Burns, 2010  Peter's Scarlet-eye  NW Costa Rica  M T1 T2 T3 S I1 I2 H G

Porphyrogenes convexus Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes spadix Austin & O. Mielke, 2008  TL: Brazil (Rd)  M OI

Porphyrogenes splendidus Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes simulator Austin & O. Mielke, 2008  TL: Bolivia  M OI

Porphyrogenes sula (R. Williams & E. Bell, 1940)  Sula Scarlet-eye  Honduras, possibly Costa Rica  M T S I G

Porphyrogenes spina Austin & O. Mielke, 2008  Spina Scarlet-eye  Panama  M T G

Porphyrogenes sporta Austin & O. Mielke, 2008  TL: Colombia  M OI

Porphyrogenes glavia Evans, 1952  Glavia Scarlet-eye  Panama  M T G

Porphyrogenes sparta Evans, 1952  TL: Brazil (Pa)  M T

Porphyrogenes spoda Evans, 1952  Spoda Scarlet-eye  Panama  M T G

Porphyrogenes stupa Evans, 1952  TL: ?  M T

Porphyrogenes passalus (Herrich-Schäffer, 1869)  Passalus Scarlet-eye  Panama, Colombia, Bolivia, Brazil (at least)  M T S G

Porphyrogenes virgatus (Mabille, 1888)  Virgatus Scarlet-eye  Honduras, Guyana, Peru  M T OI

Porphyrogenes eudemus (Mabille, 1888)  Eudemus Scarlet-eye  Panama, Peru  M T OI TI S

Porphyrogenes unassociated_unnamed_female_#1 ,  TL: not specified  M S IL

Porphyrogenes unassociated_unnamed_female_#2 ,  TL: not specified  M IL

Porphyrogenes unassociated_unnamed_female_#3 ,  Mystery Scarlet-eye #3  Panama  M S

Porphyrogenes unassociated_unnamed_female_#4 ,  Mystery Scarlet-eye #4  Panama  M S

Porphyrogenes unassociated_unnamed_female_#5 ,  Mystery Scarlet-eye #5  Costa Rica  M S

Porphyrogenes unassociated_unnamed_females ,  Mystery Female Scarlet-eyes  Porphyrogenes females whose identity remains a mystery  M S HS

Porphyrogenes unassociated_unnamed_male ,  Mystery Scarlet-eye #6  Costa Rica  M S G

Ocyba  Lindsey, 1925

Ocyba calathana (Hewitson, 1868)  Yellow-rimmed Scarlet-eye  TL: Colombia

Ocyba calathana calanus (Godman & Salvin, 1894)  Mexico to Costa Rica  M T OI S1 S2 IL L I G

Ocyba calathana calathana (Hewitson, 1868)  Panama and Colombia  M T S IL

Ocyba calathana compusa (Hewitson, 1868)  TL: Brazil (Am)  M T

Oileides  Hübner, [1825]

Oileides azines (Hewitson, 1867)  TL: Brazil (Am)  M T OI S L

Oileides fenestratus (Gmelin, [1790])  TL: [French Guiana]  M T S

Oileides vulpinus Hübner, [1825]  TL: ?

Oileides vulpinus guyanensis (Mabille & Boullet, 1912)  TL: French Guiana  M T S

Oileides vulpinus vulpinus Hübner, [1825]  TL: ?  M OI S

Oileides amazonensis (E. Bell, 1947)  TL: Colombia

Oileides amazonensis amazonensis (E. Bell, 1947)  TL: Colombia  M T S

Oileides amazonensis renta (Evans, 1952)  TL: Peru  M T

Aurina  Evans, 1937

Aurina dida Evans, 1937  TL: "Ivory Coast"  M T OI OD S G Txt

  Subfamily Pyrginae

  Tribe Pyrrhopygini

Azonax  Godman & Salvin, 1893

Azonax typhaon (Hewitson, 1877)  Typhaon Skipper  SE Mexico (OAX), Nicaragua, Colombia, Guyana  M T S L

Myscelus  Hübner, [1819]

Myscelus nobilis (Cramer, 1777)  Noble Myscelus  Surinam, Venezuela, Peru, Brazil, Bolivia  M T S HS IL

Myscelus amystis (Hewitson, 1867)  Widespread Myscelus  TL: Colombia

Myscelus amystis hages Godman & Salvin, 1893  E & W Mexico to Panama  M T S1 S2 HS L I Ha G

Myscelus amystis muttra Evans, 1951  Colombia (Valle del Cauca)  M T

Myscelus amystis amystis (Hewitson, 1867)  Trinidad, Venezuela, Colombia  M T TI S

Myscelus amystis distinctus Röber, 1925  Ecuador  M S G

Myscelus amystis mysus Evans, 1951  Peru, Bolivia  M T S

Myscelus amystis epigona Herrich-Schäffer, 1869  S Brazil, Paraguay, Argentina  M T TI S HS IL

Myscelus phoronis (Hewitson, 1867)  Phoronis Myscelus  TL: Colombia

Myscelus phoronis phoronis (Hewitson, 1867)  Guatemala, Panama, Colombia, Venezuela, Ecuador, Peru, Bolivia  M T TI S IL L

Myscelus phoronis caucanus Staudinger, 1888  Colombia  M T S

Myscelus belti Godman & Salvin, 1879  Belt's Myscelus  SE Mexico (OAX, VER, CHIS) to Venezuela, Colombia & Ecuador  M T TI S I1 I2 I3

Myscelus perissodora Dyar, 1914  Dyar's Myscelus  S Mexico (GRO, OAX, VER) to Colombia & Venezuela  M T S L I G

Myscelus pegasus Mabille, 1903  Pegasus Myscelus  French Guiana, Venezuela, Ecuador  M T TI S

Myscelus epimachia Herrich-Schäffer, 1869  Red Myscelus  TL: ?

Myscelus epimachia verina Evans, 1951  Ecuador  M T S

Myscelus epimachia epimachia Herrich-Schäffer, 1869  Venezuela, Colombia, Ecuador, Peru, Bolivia, S Brazil  M S IL

Myscelus epimachia edix Evans, 1951  SE Brazil, Paraguay  M T S

Myscelus draudti N. Riley, 1926  Draudt's Myscelus  Peru, Bolivia  M T S

Myscelus santhilarius (Latreille, [1824])  Hilarious Myscelus  SE Brazil  M T S IL

Myscelus assaricus (Cramer, 1779)  Fiery Myscelus  TL: Surinam

Myscelus assaricus assaricus (Cramer, 1779)  French Guiana, Surinam, N Brazil  M T S G

Myscelus assaricus mapirica (Strand, 1921)  S Venezuela, Peru, Brazil (Ro)  M T S IL

Myscelus assaricus michaeli Nicolay, 1975  SE Mexico (OAX, CHIS), Guatemala, Costa Rica, Panama  M T OI S1 S2 L I G

Myscelus pardalina (C. Felder & R. Felder, 1867)  Flaming Myscelus  TL: Colombia

Myscelus pardalina kesselringi O. Mielke & Casagrande, 2011  Peru  M T OI

Myscelus pardalina guarea Evans, 1951  N Brazil (Para)  M T G

Myscelus pardalina pardalina (C. Felder & R. Felder, 1867)  Colombia, Ecuador, Peru, Bolivia, Brazil  M T S IL

Myscelus pardalina yacutinga O. Mielke & Casagrande, 2011  Argentina & Paraguay  M T OI

Granila  Mabille, 1903

Granila paseas Hewitson, 1857  Paseas Skipper  SE Brazil, Paraguay  M T OI S IL

Passova  Evans, 1951

Passova nigrocephala (E. Bell, 1934)  Black-headed Firetip  Colombia  M T

Passova ganymedes (E. Bell, 1931)  Mimetic Firetip  TL: Colombia

Passova ganymedes ganymedes (E. Bell, 1931)  Colombia  M T

Passova ganymedes gulia Evans, 1951  Ecuador  M T

Passova ganymedes gelina Evans, 1951  Peru  M T S

Passova gellias (Godman & Salvin, 1893)  Gellias Firetip  Belize, Honduras, Costa Rica, Panama & Venezuela  M T S1 S2 I1 I2

Passova polemon (Hopffer, 1874)  Polemon Firetip  S Brazil  M T S L

Passova glacia Evans, 1951  Shining Firetip  Surinam, French Guiana  M S

Passova gazera Hewitson, 1866  Gazera Firetip  Brazil (Amazonas)  M T TI S

Passova greta Evans, 1951  Great Firetip  Peru, Bolivia  M T S

Passova vilna Evans, 1951  Vilna Firetip  TL: Bolivia

Passova vilna vilna Evans, 1951  Bolivia

Passova vilna vanta Evans, 1951  Bolivia  M T

Passova passova Hewitson, 1866  Passova Firetip  TL: Brazil (Am); French Guiana

Passova passova gortyna Hewitson, 1866  N Brazil  M T TI S G

Passova passova passova Hewitson, 1866  French Guiana  M T S

Passova passova rudex Evans, 1951  N Brazil, Peru  M T S

Passova passova styx Möschler, 1879  Colombia, Venezuela, Bolivia  M T S

Passova passova practa Evans, 1951  S Brazil, Paraguay  M T S

Passova passova stacer Evans, 1951  Surinam, Guyana  M T

Aspitha  Evans, 1951

Aspitha aspitha Hewitson, 1866  Aspitha Firetip  TL: Brazil (Pa/Am)

Aspitha aspitha parima Plötz, 1886  Trinidad, French Guiana, Surinam, N Brazil

Aspitha aspitha aspitha Hewitson, 1866  N Brazil (Pa, Am)  M T TI S

Aspitha aspitha rufescens (N. Riley, 1919)  Brazil (Rd)  M T

Aspitha leander (Boullet, 1912)  Leander Firetip  SE Mexico (OAX), Panama, Colombia  M T S IL G Txt

Aspitha bassleri (E. Bell, 1940)  Bassler's Firetip  Peru  M T S

Aspitha agenoria Hewitson, 1876  Red-lobed Firetip  TL: Peru

Aspitha agenoria agenoria Hewitson, 1876  Colombia, Peru, Bolivia  M T S

Aspitha agenoria sanies (H. Druce, 1908)  Peru, Bolivia  M T S IL

Aspitha agenoria cruor (H. Druce, 1908)  Peru  M T

Zonia  Evans, 1951

Zonia zonia Evans, 1951  Zonia Skipper  TL: Brazil (Am)

Zonia zonia panamensis Nicolay, 1975  Belize, Panama  M T OI S G

Zonia zonia zonia Evans, 1951  N Brazil (Am)  M T TI

Zonia zonia diabo O. Mielke & Casagrande, 1998  SE Brazil (SP)  M T

Olafia  Nemésio, 2005

Olafia roscius (Hopffer, 1874)  Olaf's Firetip  TL: Brazil

Olafia roscius iphimedia Plötz, 1886  SE Brazil (SP)  M T S IL

Olafia roscius flavomaculata (E. Bell, 1937)  SE Brazil (Pr, SC)  M T

Olafia roscius roscius (Hopffer, 1874)  SE Brazil (RS), Paraguay, Argentina  M T S

Oxynetra  C. Felder & R. Felder, 1862

Oxynetra stangelandi Grishin & Burns, 2013  Stangeland's Firetip  currently known only from ACG in Costa Rica  M T1 T2 L I1 I2

Oxynetra hopfferi Staudinger, 1888  Hopffer's Firetip  SE Mexico (VER), Costa Rica, Panama  M T OI OD TI S G

Oxynetra semihyalina C. Felder & R. Felder, 1862  Felder's Firetip  Colombia, Ecuador, Peru, Bolivia  M T OD S

Oxynetra confusa Staudinger, 1888  Confusing Firetip  Venezuela (E slope Andes), Ecuador, Peru, Bolivia, [French Guiana?]  M T S

Pyrrhopyge  Hübner, [1819]

Pyrrhopyge zenodorus Godman & Salvin, 1893  Red-headed Firetip  SE Mexico to Costa Rica  M T S L I1 I2 G

Pyrrhopyge evansi E. Bell, 1947  Evans' Firetip  TL: Colombia

Pyrrhopyge evansi evansi E. Bell, 1947  Costa Rica, Panama, Colombia, Venezuela (Bolivar), French Guiana, [Ecuador?]  M T S G

Pyrrhopyge evansi borburata Orellana, [2010]  Venezuela (N slope Cordillera de la Costa, Carabobo)

Pyrrhopyge phidias (Linnaeus, 1758)  Original Firetip  TL: Asia

Pyrrhopyge phidias hyperici Hübner, 1823  Brazil (Para)

Pyrrhopyge phidias latifasciata A. Butler, 1873  Colombia  M T

Pyrrhopyge phidias bixae (Linnaeus, 1758)  Surinam, French Guiana, N Brazil to Matto Grosso

Pyrrhopyge phidias rusca Evans, 1951  Ecuador, Peru, SW Brazil  M T

Pyrrhopyge phidias leucoloma Erschoff, 1875  Peru, Bolivia

Pyrrhopyge phidias garata Hewitson, 1866  Guyana, Surinam, [N Venezuela?]  M T

Pyrrhopyge phidias phidias (Linnaeus, 1758)  Trinidad, Surinam, Guyana, Venezuela, Colombia, Peru, Bolivia, Paraguay, Brazil  M T S

Pyrrhopyge phidias guianae E. Bell, 1932  Trinidad, Guyana, French Guiana  M T

Pyrrhopyge aziza Hewitson, 1866  Aziza Firetip  TL: Colombia

Pyrrhopyge aziza aziza Hewitson, 1866  Colombia  M T TI

Pyrrhopyge aziza lexos Evans, 1951  Guyana, Colombia, Ecuador, Peru  M T S

Pyrrhopyge aziza araethyrea Hewitson, 1870  Ecuador  M T TI S

Pyrrhopyge aziza troja Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge aziza arbor Evans, 1951  Colombia, Venezuela  M T S

Pyrrhopyge aziza attis E. Bell, 1931  Bolivia  M T

Pyrrhopyge aziza subnubilus Hayward, 1935  Argentina

Pyrrhopyge pusca Evans, 1951  Pusca Firetip  Peru, Bolivia  M T S

Pyrrhopyge proculus Hopffer, 1874  Proculus Firetip  TL: Guyana

Pyrrhopyge proculus lina E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge proculus cardus Mabille, 1891  N Brazil  M T

Pyrrhopyge proculus hyleus Mabille, 1891  N Brazil  M T

Pyrrhopyge proculus cintra Evans, 1951  French Guiana, Surinam, Guyana, N Brazil, NE Peru  M T S

Pyrrhopyge proculus draudti E. Bell, 1931  Venezuela, Colombia, Ecuador, Peru, Bolivia (Andean)  M T

Pyrrhopyge proculus proculus Hopffer, 1874  Trinidad, Guyana, Venezuela (E Cordillera de la Costa), Colombia, N Brazil  M T

Pyrrhopyge infantilis H. Druce, 1908  Infant Firetip  TL: Peru

Pyrrhopyge infantilis infantilis H. Druce, 1908  Peru, Bolivia  M T S

Pyrrhopyge infantilis agala Evans, 1951  Bolivia  M T S

Pyrrhopyge thericles Mabille, 1891  Impostor Firetip  TL: Brazil (Am)

Pyrrhopyge thericles orientis E. Bell, 1947  N Brazil (Para)  M T

Pyrrhopyge thericles thericles Mabille, 1891  NE Brazil  M T

Pyrrhopyge thericles ponicia Evans, 1951  French Guiana, Surinam, N Brazil  M T S

Pyrrhopyge thericles pseudophidias E. Bell, 1931  Costa Rica, Panama, Venezuela, Colombia, Ecuador, Peru, N Brazil  M T S

Pyrrhopyge thericles raymondi Orellana, [2010]  Venezuela (Cordillera de la Costa, Carabobo, Aragua, Guarico, Miranda)

Pyrrhopyge thericles fola Evans, 1951  Colombia, SW Venezuela  M T

Pyrrhopyge thericles rileyi E. Bell, 1931  Ecuador, Bolivia  M T

Pyrrhopyge thericles grinda Evans, 1953  Guyana  M T

Pyrrhopyge thericles ronda Evans, 1953  Trinidad  M T

Pyrrhopyge amythaon E. Bell, 1931  Amythaon Firetip  TL: Brazil (Pe)

Pyrrhopyge amythaon gradens Evans, 1951  N Brazil (Para)  M T S

Pyrrhopyge amythaon peron E. Bell, 1947  Venezuela (Bolivar), Colombia, NE Peru  M T S

Pyrrhopyge amythaon polka Evans, 1951  French Guiana  M T S

Pyrrhopyge amythaon perula Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge amythaon orino Evans, 1951  Colombia, Venezuela (S of Orinoco), Guyana  M T S

Pyrrhopyge amythaon pollio Evans, 1951  Peru  M T

Pyrrhopyge amythaon podina Evans, 1951  Peru, Bolivia  M T

Pyrrhopyge amythaon amythaon E. Bell, 1931  Brazil (Pernambuco)  M T

Pyrrhopyge sergius Hopffer, 1874  Sergius Firetip  TL: Peru

Pyrrhopyge sergius selina Evans, 1951  Brazil (Para)  M T S

Pyrrhopyge sergius andros Evans, 1951  Colombia, Venezuela (Amazonas)  M T

Pyrrhopyge sergius andronicus E. Bell, 1931  Ecuador, Peru, Bolivia  M T

Pyrrhopyge sergius semana Evans, 1951  Surinam, French Guiana  M T S

Pyrrhopyge sergius sergius Hopffer, 1874  Peru  M T S

Pyrrhopyge sergius ganus E. Bell, 1947  Guyana, Venezuela (Cordillera de la Costa), [Colombia, Ecuador, Peru, Brazil?]  M T

Pyrrhopyge sergius josephina Draudt, 1921  Peru, Bolivia  M T

Pyrrhopyge caribe Orellana, [2010]  Caribbean Firetip  TL: Venezuela

Pyrrhopyge caribe caribe Orellana, [2010]  Venezuela (Aragua, Carabobo, Dtto. Capital, Lara, Trujillo, Vargas)

Pyrrhopyge caribe camachoi Orellana, [2010]  Venezuela (Gran Sabana)

Pyrrhopyge erazoae Orellana, [2010]  Erazo's Firetip  Venezuela (Bolivar)  M S

Pyrrhopyge charybdis Westwood, 1852  Charybdis Firetip  TL: [Brazil (MG)]

Pyrrhopyge charybdis semita Evans, 1951  Bolivia, S Brazil  M T S L

Pyrrhopyge charybdis charybdis Westwood, 1852  S Brazil  M T OI L

Pyrrhopyge cressoni E. Bell, 1932  Cresson's Firetip  Ecuador, Bolivia  M T S

Pyrrhopyge pelota Plötz, 1879  Pelota Firetip  Bolivia, Paraguay, S Brazil  M T

Pyrrhopyge amyclas (Cramer, 1779)  Yellow-edged Firetip  TL: Surinam

Pyrrhopyge amyclas amyclas (Cramer, 1779)  Trinidad, Venezuela, Guyana, Surinam, French Guiana, Brazil

Pyrrhopyge amyclas denticulata Herrich-Schäffer, 1869  Ecuador, Peru, Brazil  M S

Pyrrhopyge phylleia Hewitson, 1874  Orange-edged Firetip  TL: Bolivia

Pyrrhopyge phylleia delos Evans, 1951  Peru  M T S

Pyrrhopyge phylleia phylleia Hewitson, 1874  S Peru, Bolivia  M T TI S L G

Pyrrhopyge arax Evans, 1951  Arax Firetip  SE Peru, Bolivia  M T S

Pyrrhopyge haemon Godman & Salvin, 1893  Tawny-rimmed Firetip  Costa Rica  M T S

Pyrrhopyge punctata Röber, 1925  Songo Firetip  Bolivia (Rio Songo)

Pyrrhopyge papius Hopffer, 1874  Papius Firetip  TL: Colombia; Bolivia

Pyrrhopyge papius pasca Evans, 1951  Colombia  M T S

Pyrrhopyge papius papius Hopffer, 1874  Venezuela, Colombia, Ecuador, Peru (Andean)  M T S

Pyrrhopyge frona Evans, 1951  Inca Firetip  Peru  M T S

Pyrrhopyge melanomerus Mabille & Boullet, 1908  Melanomerus Firetip  TL: Bolivia

Pyrrhopyge melanomerus patma Evans, 1951  S Ecuador, NE Peru  M T

Pyrrhopyge melanomerus melanomerus Mabille & Boullet, 1908  Peru, Bolivia  M T L

Pyrrhopyge decipiens Mabille, 1903  Red-spotted Firetip  [Ecuador?], Peru, Bolivia  M T S

Pyrrhopyge placeta Evans, 1951  Placeta Firetip  Brazil (ES)  M T

Pyrrhopyge hadassa Hewitson, 1866  Hadassa Firetip  TL: [Ecuador]

Pyrrhopyge hadassa hadassa Hewitson, 1866  Colombia, Ecuador, Peru  M T TI S

Pyrrhopyge hadassa henna Evans, 1951  Peru  M T

Pyrrhopyge hadassa hanga Evans, 1951  Peru  M T

Pyrrhopyge hadassa pseudohadassa Mabille & Boullet, 1908  Peru  M T S L

Pyrrhopyge hadassa halma Evans, 1951  Peru, Bolivia  M T S

Pyrrhopyge terra Evans, 1951  Terra Firetip  Bolivia  M T S

Pyrrhopyge telassina Staudinger, 1888  Telassina Firetip  TL: Peru; Bolivia

Pyrrhopyge telassina tagra Evans, 1951  Peru  M T S

Pyrrhopyge telassina telassina Staudinger, 1888  Peru  M T S

Pyrrhopyge telassina shiva Evans, 1951  SE Peru, Bolivia  M T L

Pyrrhopyge telassa Hewitson, 1866  Telassa Firetip  TL: [Ecuador]

Pyrrhopyge telassa telassa Hewitson, 1866  Ecuador  M T TI IL

Pyrrhopyge telassa phaeax Hopffer, 1874  Peru  M T S L

Pyrrhopyge telassa silex Evans, 1951  Peru  M T

Pyrrhopyge telassa croceimargo Mabille & Boullet, 1908  S Peru, Bolivia  M T S HS L

Pyrrhopyge martena Hewitson, [1869]  Martena Firetip  Ecuador  M T OI

Pyrrhopyge schausi E. Bell, 1931  Schaus' Firetip  Colombia, Ecuador, N Peru  M T S

Pyrrhopyge sadia Evans, 1951  Sadia Firetip  Ecuador, Peru  M T S

Pyrrhopyge creona H. Druce, 1874  Creona Firetip  Peru, Brazil (Am)  M T S

Pyrrhopyge kelita Hewitson, [1869]  Streaked Firetip  SE Peru, Bolivia  M T OI

Pyrrhopyge crista Evans, 1951  Crista Firetip  Bolivia  M T

Pyrrhopyge crida Hewitson, 1871  White-banded Firetip  SE Mexico to Colombia and Ecuador  M T TI S1 S2 IL I1 I2 G

Pyrrhopyge mopsus (E. Bell, 1931)  Mopsus Firetip  Peru  M T S

Pyrrhopyge sarpedon (E. Bell, 1931)  Sarpedon Firetip  Peru (Amazonas)  M T

Pyrrhopyge arinas (Cramer, 1777)  Arinas Firetip  TL: Surinam

Pyrrhopyge arinas arinas (Cramer, 1777)  French Guiana, Surinam, N Brazil, NE Peru  M T TI S IL

Pyrrhopyge arinas temenos (E. Bell, 1931)  Peru  M T S

Pyrrhopyge creusae (E. Bell, 1931)  Creusae Firetip  French Guiana  M T

Pyrrhopyge tatei (E. Bell, 1932)  Tate's Firetip  Venezuela  M T

Pyrrhopyge boulleti Le Cerf, 1922  Boullet's Firetip  Colombia, Venezuela (Tachira)  M S

Gunayan  O. Mielke, 2002

Gunayan rhacia Hewitson, [1875]  Rhacia Skipper  S Brazil  M T OI

Gunayan rubricollis (Sepp, [1841])  Red-collared Skipper  French Guiana, Peru, Bolivia, Brazil  M T TI S IL L

Gunayan timaeus (E. Bell, 1931)  Timaeus Skipper  Colombia, Peru  M T S

Yanguna  E. Watson, 1893

Yanguna tetricus E. Bell, 1931  Tetricus Skipper  Guyana, Peru  M T

Yanguna cometes (Cramer, 1779)  Comet Skipper  TL: Surinam

Yanguna cometes cometes (Cramer, 1779)  French Guiana, N Brazil (Para)

Yanguna cometes staudingeri Plötz, 1879  Surinam, Venezuela (Bolivar), S Colombia (upper Putumayo), Ecuador, N Peru  M S IL L

Yanguna cometes angeli Orellana, [2010]  Venezuela (Tachira)

Yanguna cometes cometides Mabille & Boullet, 1908  Peru, Bolivia  M S IL

Yanguna thelersa Hewitson, 1866  Thelersa Skipper  French Guiana, N Brazil, Ecuador, Peru  M T OI S

Yanguna cosyra H. Druce, 1875  Cosyra Skipper  Guatemala to Colombia and Ecuador  M T S1 S2 IL I1 I2 I3 G

Yanguna spatiosa Hewitson, 1870  Spatiosa Skipper  TL: Ecuador

Yanguna spatiosa spatiosa Hewitson, 1870  W Colombia, Ecuador  M T TI S IL L

Yanguna spatiosa mabillei H. Druce, 1909  Peru  M T S L