Chlosyne palla flavula (W. Barnes & McDunnough, 1918)

Chlosyne palla flavula (W. Barnes & McDunnough, 1918)
(Northern Checkerspot)

Distribution and Larval Foodplants:

Asteraceae: Chrysothamnus viscidiflorus (Hook.) Nutt., Eucephalus engelmannii (D.C. Eat.) Greene

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Chlosyne Butler, 1870

Cist. Entomol. 1(3): 38, no. 37. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist., 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). The synonymy for this genus follows Wahlberg and Zimmerman (2000), Cladistics 16(4): 347-363.

= ‡Morpheis Geyer, [1833]

In: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett., 3: pl [5], figs. 1-4, by monotypy. Preoccupied by Morpheis Hübner, [1820], Verz. bekannt. Schmett., 13: 196.

= ‡Synchloe E. Doubleday, [1845]

List Spec. Lepid. Brit. Mus. 1: 76. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 141, no. 253. Preoccupied by Synchloe Hübner, 1818, Zutr. Samml. exot. Schmett. 1: 26.

= ‡Coatlantona W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 178. Type-species: Papilio janais Drury, 1782, Ill. Nat. Hist. 3: 22, index, pl. 17, figs. 5, 6, as replacement name. Proposed to replace Synchloe E. Doubleday, 1844, preoccupied (Code Articles 60.3, 67.8). A junior objective synonym of Chlosyne Butler, 1870.

= Anemeca W. F. Kirby, 1871

Syn. Cat. diurn. Lepid.: 179. Type-species: Morpheis ehrenbergii Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, as replacement name. Proposed to replace Morpheis Geyer, [1833], in: Hübner, Samml. exot. Schmett. 3: pl [5], figs. 1-4, preoccupied (Code Articles 60.3, 67.8).

= Charidryas Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea nycteis E. Doubleday, 1847, in: Gen. diurn. Lepid. (1)(12): pl. 23, fig. 3; (21): 181, no. 23 (1848), by original designation.

= ‡Limnaecia Scudder, 1872

4th Ann. Rept. Peabody Acad. Sci. (1871): 47. Type-species: Melitaea harrisii Scudder, 1863, Proc. Essex Inst. 3: 167-168, no. 36, by original designation. Preoccupied by Limnaecia Stainton, 1851, Suppl. Cat. Brit. Pteroph.: 4.

= Thessalia Scudder, 1875

Bull. Buffalo Soc. Nat. Sci. 2(4): 238, 265, no. 49. Type-species: Melitaea leanira C. Felder & R. Felder, 1860, Wien. Entomol. Montaschr. 4(4): 106, no. 64, by original designation.

Chlosyne palla (Boisduval, 1852)

Annls. Soc. Entomol. Fr. (2)10(2): 305, no. 48; figured by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 259, figs. 2173 ♂ D&V, 2174 ♀ D&V.

Original Combination: Melitaea palla

Type Locality: “Dans une grande partie de la Californie”; defined as “vic. San Francisco, California” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 156; corrected to “Hwy. 70 at Chambers Creek, ca. 6 rd. Mi SW of Belden, North Fork Feather River Canyon, 1850' elev., Plumas County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 18, figs. 99 ♂ D, 100 ♂ V.

= ‡”wardi” (Oberthür, 1914)

Étud. Lépid. Comp. 9(2): 79, pl. 259, fig. 2175 ♂ D&V (“holotype”).

Original Combination: [Melitaea Palla] aberration Wardi

Type Locality: Not stated; “Californie” implied from the title of the paper.

Types: “Holotype” in USNM; figured by J. A. Comstock (1927), Butts. Calif.: pl. 27, fig. 2 ♂ D&V.

= ‡”stygiana” (J. A. Comstock, 1926)

Bull. South. Calif. Acad. Sci. 25(1): 32; figured by J. A. Comstock (1927), Butts. Calif.: pl. 36, fig. 19 ♀ D (“holotype”).

Original Combination: Melitaea palla ab. stygiana

Type Locality: “Willow Ranch, Modoc County, Calif.” [California]

Types: “Holotype” in LACM.

Chlosyne palla flavula (W. Barnes & McDunnough, 1918)

Contr. Nat. Hist. Lepid. N. Am. 4(2): 73-74, pl. 11, figs. 14 ♂ D, 15 ♂ V.

Original Combination: Melitaea flavula

Type Locality: “Glenwood Spgs., Colo....Hall Valley, Colo.”; defined as “Glenwood Springs, Colorado” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 157. [Garfield County]

Types: Syntypes in USNM; one figured by W. Holland (1931), Butt. Book (rev. ed.): pl. 56, fig. 25 ♂ D.

Bibliography

Barnes, William and James Halliday McDunnough. 1918. Notes and new species. Contributions to the natural History of the Lepidoptera of North America 4(2): 61-208 + [4]; pls. 11-25 {15 May 1918} [description of Melitaea flavula]

Brown, Frederick Martin. 1957. Colorado butterflies [Assisted by Donald Eff and Bernard Rotger.] Denver, Denver Museum Natural History: vii + 368 pp.; 314 unnumbered figs. {1957}

Ferris, Clifford Duras and Michael S. Fisher. [1978]. Charidryas flavula Barnes & McDunnough (Nymphalidae): A question of identity. Journal of Research on the Lepidoptera 16(3): 133-140, 15 figs. {Sep 1977, [May 1978]}

Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figures [contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford] {1981}

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960}

Holland, William Jacob. 1931. The butterfly book. New and thoroughly revised edition. A popular and scientific manual, describing and depicting all the butterflies of the United States and Canada. Garden City, New York; Doubleday, Doran & Co. xii + 424 pp., 77 pls., 198 + [1] text figures. {1931}

Miller, Lee Denmar and Frederick Martin Brown. 1981. A catalogue/checklist of the butterflies of America north of Mexico. Memoir. The Lepidopterists= Society (2): vii + 280 pp. {1981}

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