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Colias p. philodice Godart, 1819
(Clouded Sulphur)

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Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Colias [Fabricius], 1807

In: Illiger, Mag. f. Insektenk. 6: 284, no. 24. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by designation of I.C.Z.N. Opinion 146.

      =    ‡Eurymus Horsfield, [1829]

Descr. Cat. Lepid. Ins. Mus. E. India Co. (2): 129, 134. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation. Preoccupied by Eurymus Rafinesque, 1815, Analyse Nat.: 117, and a junior objective synonym of Colias.

=    Scalidoneura Butler, 1871

Proc. Zool. Soc. Lond. 1871(1): 250-251. Type-species: Scalidoneura hermina Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 251, pl. 19, fig. 5, by original designation.

=    Eriocolias J. Watson, 1895

Entomol. 28(385): 167. Type-species: Papilio edusa Fabricius, 1787, Mantissa Ins. 2: 23, no. 240, by original designation (= Papilio croceus Geoffroy (1875), in: Fourcroy, Entomol. Paris 2: 250, no. 48).

      =    ‡Coliastes Hemming, 1931

Entomol. 64(12)[823]: 273. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation; a junior objective synonym of Colias.

=    Mesocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias vauthierii Guérin-Méneville, [1830], in: Duperrey, Voy. Coquille, 1(Zool.)Atlas(18): pl. Ins. 15, fig. 2, by original designation.

=    Protocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias imperialis Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 250, pl. 19, fig. 2, by original designation (= Colias ponteni Wallengren, 1860, Wien. entomol. Monatschr. 4(2): 33, no.1; synonym).

=    Palaeocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21-22, 32. Type-species: Colias ponteni Wallengren, 1860, Wien. Entomol. Montaschr. 4(2): 33, by original designation.

=    Neocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21, 32. Type-species: Papilio erate Esper, 1805, Die Schmett. 1(suppl.)(2)(11): 13-14, pl. 119 (Cont. 74), fig. 3, by original designation.

=    Eucolias Berger, 1986

Lambillionea 86(7/8, suppl.): 22, 33. Type-species: Papilio palaeno Linnaeus, 1761, Fauna Svecica (ed. 2): 272, no. 1041, by original designation.

=    Similicolias Berger, 1986

Lambillionea 86(7/8, suppl.): 23, 33. Type-species: Papilio lesbia Fabricius, 1775, Syst. Entomol.: 477, no. 149, by original designation.

=    Paracolias Berger, 1986

Lambillionea 86(7/8, suppl.): 24, 34. Type-species: Colias dimera E. Doubleday, [1847], in: Gen. diurn. Lepid. (1): pl. 9, fig. 3; (7): 74, no. 15, by original designation.

=    Asiocolias Korb, 2005

Cat. Butt. ex USSR: 20. Type-species: Colias christophi Grum-Grshimaïlo, 1885, in: Romanov, Mém. Lépid. 2: 220, by original designation.

Ganura Zetterstedt, [1839]

Ins. Lappon. (5): col. 903. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71. This name was published in synonymy and is unavailable (Code Article 11.6).

 

Colias philodice Godart, 1819

 

Colias philodice philodice Godart, 1819

Ency. méth. 9(Ins.)(1): (87), 100-101, no. 35.

Original Combination: Colias Philodice

Type Locality: “l'Amérique septentrionale, particulièrement dans la Virginie”; defined as “Virginia” by Klots (1951), Field Guide Butts. N. Am.: 185.

Types: Location of type not known.

                   =    dorippe Godart, 1819

Ency. méth. 9(Ins.)(1): (87), 101, no. 36; cited Papilio palaeno (sensu Cramer (1775), Uitl. Kapellen 1(2): pl. 14, figs. F, G).

Original Combination: Colias Dorippe

Type Locality: “Cap de Bonne-Espérance” in error. [Cape Good Hope, South Africa]

Types: Type(s) may be lost.

                   =    anthyale (Hübner, 1823)

Samml. exot. Schmett. 2: pl. [54], figs. 307 ♂ D, 308 ♂ V [1821]; (text): 21, no. 154 (1832, name given).

Original Combination: Zerene Anthyale

Type Locality: “Pennsylvanien” [Pennsylvania]

Types: Type(s) probably lost.

                   =    santes Fitch, 1854

Trans. N. Y. State Agric. Soc. 13(1853): 378.

Original Combination: Colias santes

Type Locality: “New York”; J. Barnes (1988), Bull. New York St. Mus. (461): 112, no. 334, noted that the original specimen (#6680) was labeled “Salem, NY.” [Washington County, New York]

Types: Probably lost; J. Barnes (1988), ibid., noted that there were no extant specimens in the NYSM.

                   =    tegea Sievers, 1859

Etud. Entomol. (Motschulsky) 8: 146.

Original Combination: Colias Tegea

Type Locality: “environs de New York”

Types: Location of type(s) not known.

                         =    ‡notatus (Megerle, 1803)

Cat. Ins. Viennae Austriae 28 Novembris 1803, auctionis: [81], no. 436.

Type Locality: “Georgia”

Original Combination: [Papilio] notatus

Types: Probably none designated.

I.C.Z.N. Opinion 1710 placed this work on the Official Index of Rejected and Invalid Works in Zoological Nomenclature. This name has at times been credited to A. Clark and L. Clark (1941), Proc. Entomol. Soc. Wash. 43(4): 83, no. 1; their mention of the name represents a nomen nudum.

                         =    ‡”alba” Strecker, 1878

Butts. Butts. & Moths N. Am., Comp. Syn. Cat.: 82, no. 58a; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 2, fig. 6 ♀ D, pl. Colias III, figs. 5 ♀ D, 6 ♀ V.

Original Combination: [Colias Philodice] ab. a. ♀ Alba

Type Locality: Not stated.

Types: “Type(s)” may be in CMNH.

                         =    ‡”nig” Strecker, 1878

Butts. & Moths N. Am., Comp. Syn. Cat.: 82, no. 58b; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 3, figs. 8 ♂ D, 9 ♂ V.

Original Combination: [Colias Philodice] ab. b. ♂ Nig

Type Locality: “near Palmyra, New York...Montreal, Canada”; “lectotype” from “Montreal.” [Quebec]

Types: The specimen from Montreal in FMNH is the sole remaining “syntype”; L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 220, note 281, designated this as “lectotype.”

                         =    ‡”virida” Strecker, 1878

Butts. & Moths N. Am., Comp. Syn. Cat.: 82, no. 58c.

Original Combination: [Colias Philodice] ab. c. ♀ Virida

Type Locality: “taken at the same time and place as the preceding [nig]” “near Montreal, Canada” [Quebec]

Types: “Holotype” in FMNH.

                         =    ‡”hybrida” Strecker, 1878

Butts. & Moths N. Am., Comp. Syn. Cat.: 82, no. 58d; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 3, fig. 1 ♂ D.

Original Combination: [Colias Philodice] ab. d. ♀ Hybrida

Type Locality: “Illinois, Georgia”; determined to be “Rock Island, Illinois” from Edwards’ figure. [Rock Island County]

Types: “Holotype” is the specimen figured by W. H. Edwards (1876), ibid., possibly in CMNH.

                         =    ‡”albinic” W. H. Edwards, [1885]

Trans. Am. Entomol. Soc. 11(3/4): 264, under no. 68; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 2, fig. 6 ♀ D.

Original Combination: [Colias Philodice] var. Albinic

It is doubtful that this can be considered the description of a new taxon but is included for the sake of completeness.

                         =    ‡”melanic” W. H. Edwards, [1885]

Trans. Am. Entomol. Soc. 11(3/4): 264, under no. 68; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 3, figs. 8 ♀ D, 9 ♀ V.

Original Combination: [Colias Philodice] var. Melanic

It is doubtful that this can be considered the description of a new taxon but is included for the sake of completeness.

                         =    ‡”maria” W. H. Edwards, 1885

Papilio 4(9-10): 171.

Original Combination: C[olias]. [philodice] maria

Type Locality: Not stated.

Types: “Holotype” lost.

This name represents a cyanide-stained specimen, inadvertently described.

                         =    ‡”nigridice” Scudder, 1889

Butts. E. U. S. & Can. 2(8): 1114; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 3, figs. 8 ♂ D, 9 ♂ V.

Original Combination: Eurymus phil[odice]. [♂ form] nigridice

Type Locality: “Palmyra, N. Y.” [Wayne County, New York]

Types: “Holotype” is the specimen figured by W. H. Edwards (1876), ibid., possibly in CMNH.

                         =    ‡”miscidice” Scudder, 1889

Butts. E. U. S. & Can. 2(8): 1115.

Original Combination: E[urymus]. p[hilodice]. [♀ form] miscidice

Type Locality: “Lynn, Mass.” [Essex County, Massachusetts]

Types: “Holotype” in Sanborn collection, location not known.

                         =    ‡”pallidice” Scudder, 1889

Butts. E. U. S. & Can. 2(8): 1115; 3(8): pl. 7, fig. 6 ♀ D (“holotype”); cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 2, fig. 6 ♀ D; pl. Colias III, figs. 5 ♀ D, 6 ♀ V.

Original Combination: E[urymus]. p[hilodice]. [♀ form] pallidice

Type Locality: “New England” implied from the title of the work.

Types: Location of “syntypes” not known, may be at MCZ.

                         =    ‡”suffusa” Cockerell, 1889

Entomol. 22(310): 55.

Original Combination: C[olias]. philodice suffusa

Type Locality: “Massachusetts”

Types: “Type” figured by Maynard (1886), Butts. N. England: pl. vii, fig. 57c, may be based on a specimen in the MCZ.

The text clearly indicates that this taxon was described as an individual variant.

                         =    ‡”alba” Ehrmann, 1890

Entomol. News 1(6): 93.

Original Combination: Colias philodice, Ab. alba

Type Locality: “Sarver Station, Butler County, Pa.” [Pennsylvania]

Types: Type either unrecognizable or lost.

Redescribed by Ehrmann (1890) in Entomol. News 1(8): 130.

                         =    ‡”luteitincta” Wolcott, 1893

Can. Entomol. 25(4): 104, under no. 50.

Original Combination: Colias philodice, var. luteitincta

Type Locality: “Batavia, Ills....Grand Rapids” [Kane County, Illinois; Kent County, Michigan]

Types: Type may be in USNM.

The text clearly indicates that this taxon was described as an individual variant.

                         =    ‡”albinic” Skinner, 1898

Syn. Cat. N. Am. Rhop.: 68, no. 415; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 2, fig. 6 ♀ D.

Original Combination: [Colias Philodice] var. albinic

It is doubtful that this can be considered the description of a new taxon. It has been listed in checklists and catalogues (e.g., L. Miller and F. Brown (1981), Mem. Lepid. Soc. 2: 78), but is included for the sake of completeness.

                         =    ‡”melanic” Skinner, 1898

Syn. Cat. N. Am. Rhop.: 69, no. 415; cited W. H. Edwards (1876), Butts. N. Am. 2(5): pl. Colias 3, figs. 8 ♂ D, 9 ♂ V.

Original Combination: [Colias Philodice] var. melanic

It is doubtful that this can be considered the description of a new taxon. It has been listed in checklists and catalogues (e.g., L. Miller and F. Brown (1981), Mem. Lepid. Soc. 2: 78), but is included for the sake of completeness.

                         =    ‡”nigrina” Strecker, 1900

Lepid. Rhop. Het. (suppl. 3): 19; cited Strecker (1878), Butts. & Moths N. Am., Comp. Syn. Cat.: 82, no. 58b.

Original Combination: Colias philodice ♂ form nigrina

Type Locality: “Bethlehem, Pa...Montreal, Can...Orillia, Can.” [Lehigh and Northhampton counties, Pennsylvania; Quebec and Ontario]

Types: “Syntypes” in FMNH.

This amounts to a re-description of Colias philodice ab. ♂ nig which was not cited by name.

                         =    ‡”alba” Röber, 1909

In: Seitz, Grossschmett. Erde 5(29): 91.

Original Combination: [Colias philodice] ab. alba

Type Locality: Not stated; “lectotype” from “Philadelphia.” [Philadelphia County, Pennsylvania]

Types: “Lectotype” in ZMHB, designated by Lamas (1995), Shilap Revta. Lepid. 23(91): 344.

                         =    ‡”nigrofasciata” Reiff, 1917

Lepidopterist 1(3): 22-23, pl. 3 ♀ D.

Original Combination: Colias philodice f[orm] nigrofasciata

Type Locality: “Belchertown, Massachusetts” [Hampshire County]

Types: “Holotype” in USNM.

                         =    ‡”rothkei” Reiff, 1917

Lepidopterist 1(11): 84, pl. 7 ♂ D (“holotype”).

Original Combination: Colias philodice ab. rothkei

Type Locality: “Susquehanna Valley, Pa.” [Pennsylvania]

Types: “Holotype” in AMNH.

                         =    ‡”plicaduta” Nakahara, 1926

Bull. Brooklyn Entomol. Soc. 20(5): 222-223.

Original Combination: Colias philodice f. plicaduta

Type Locality: “Ithaca, N. Y.” [Tompkins County, New York]

Types: “Holotype” in USNM.

                         =    ‡”inversata” Nakahara, 1926

Bull. Brooklyn Entomol. Soc. 20(5): 223.

Original Combination: Colias philodice ab. inversata

Type Locality: “Midvale, Allegheny Co., Pa.” [Pennsylvania]

Types: “Holotype” in USNM.

                         =    ‡”minor” F. Chermock, 1927

Bull. Brooklyn Entomol. Soc. 22(2): 118.

Original Combination: Colias philodice form plicaduta ab. minor

Type Locality: “N. S. Pittsburgh, Pa.” [Allegheny County, Pennyslvania]

Types: “Holotype” in MGCL.

                         =    ‡”ehrmanni” F. Chermock, 1927

Bull. Brooklyn Entomol. Soc. 22(2): 118.

Original Combination: Colias philodice from [sic!] plicaduta ab. Ehrmanni

Type Locality: “N. S. Pittsburgh, Pa.” [Allegheny County, Pennyslvania]

Types: “Holotype” in MGCL.

                         =    ‡”alba” F. Chermock, 1927

Bull. Brooklyn Entomol. Soc. 22(2): 119.

Original Combination: Colias philodice from [sic!] plicaduta var. alba

Type Locality: “N. S. Pittsburgh, Pa.” [Allegheny County, Pennyslvania]

Types: “Holotype” in MGCL.

                         =    ‡”albida” F. Chermock, 1928

Bull. Brooklyn Entomol. Soc. 23(4): 173.

Original Combination: Colias philodice f. plicaduta var. albida

Proposed as a replacement name for alba F. Chermock, 1927, but that name is infrasubspecific and therefore not subject to homonymy.

                         =    ‡”raritus” (Gunder, 1928)

Can. Entomol. 60(7): 163, pl. A, figs. 2 ♀ D, 2a ♀ V (“holotype”).

Original Combination: Eurymus philodice tr. f. raritus

Type Locality: “Scranton, Pa.” [Lackawanna County, Pennsylvania]

Types: “Holotype” in AMNH.

                         =    ‡”serrata” (F. Chermock, 1929)

Bull. Brooklyn Entomol. Soc. 24(1): 21.

Original Combination: Eurymus philodice var. serrata

Type Locality: “Rossgrove, near Aspinwall, Pa.” [Allegheny County, Pennsylvania]

Types: “Holotype” in MGCL.

                         =    ‡”minor” Dufrane, 1947

Bull. Annls. Soc. Entomol. Belg. 83(1/2): 69.

Original Combination: [Colias philodice f. anthyale] minor

Type Locality: “Temple, Pa. E. U. A.” [Berks County, Pennsylvania]

Types: “Syntype” in IRSN.

Dufrane probably used the name minor as a qualifier for a small specimen.

                         =    ‡”reducta” Dufrane, 1947

Bull. Annls. Soc. Entomol. Belg. 83(1/2): 69.

Original Combination: [Colias philodice f. anthyale] ab. reducta

Type Locality: “Temple, Pa. E. U. A.” [Berks County, Pennsylvania]

Types: “Syntype” in IRSN.

‡”alba” Pilate, 1882

Papilio 2(5): 65.

Original Combination: [Colias] Philodice ♀ var. Alba

Proposed without a description or indication; a nomen nudum

‡”vernalis” Scott, 1986

Butts. N. Am.: 198, no. 38.

Original Combination: [Colias philodice] form vernalis

Proposed as a “form” with no nomenclatural standing; a nomen nudum.

Bibliography

 

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