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Colias palaeno chippewa W. H. Edwards, 1870
(Palaeno Sulphur)

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Synonymy

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Colias [Fabricius], 1807

In: Illiger, Mag. f. Insektenk. 6: 284, no. 24. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by designation of I.C.Z.N. Opinion 146.

      =    ‡Eurymus Horsfield, [1829]

Descr. Cat. Lepid. Ins. Mus. E. India Co. (2): 129, 134. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation. Preoccupied by Eurymus Rafinesque, 1815, Analyse Nat.: 117, and a junior objective synonym of Colias.

=    Scalidoneura Butler, 1871

Proc. Zool. Soc. Lond. 1871(1): 250-251. Type-species: Scalidoneura hermina Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 251, pl. 19, fig. 5, by original designation.

=    Eriocolias J. Watson, 1895

Entomol. 28(385): 167. Type-species: Papilio edusa Fabricius, 1787, Mantissa Ins. 2: 23, no. 240, by original designation (= Papilio croceus Geoffroy (1875), in: Fourcroy, Entomol. Paris 2: 250, no. 48).

      =    ‡Coliastes Hemming, 1931

Entomol. 64(12)[823]: 273. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation; a junior objective synonym of Colias.

=    Mesocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias vauthierii Guérin-Méneville, [1830], in: Duperrey, Voy. Coquille, 1(Zool.)Atlas(18): pl. Ins. 15, fig. 2, by original designation.

=    Protocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias imperialis Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 250, pl. 19, fig. 2, by original designation (= Colias ponteni Wallengren, 1860, Wien. entomol. Monatschr. 4(2): 33, no.1; synonym).

=    Palaeocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21-22, 32. Type-species: Colias ponteni Wallengren, 1860, Wien. Entomol. Montaschr. 4(2): 33, by original designation.

=    Neocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21, 32. Type-species: Papilio erate Esper, 1805, Die Schmett. 1(suppl.)(2)(11): 13-14, pl. 119 (Cont. 74), fig. 3, by original designation.

=    Eucolias Berger, 1986

Lambillionea 86(7/8, suppl.): 22, 33. Type-species: Papilio palaeno Linnaeus, 1761, Fauna Svecica (ed. 2): 272, no. 1041, by original designation.

=    Similicolias Berger, 1986

Lambillionea 86(7/8, suppl.): 23, 33. Type-species: Papilio lesbia Fabricius, 1775, Syst. Entomol.: 477, no. 149, by original designation.

=    Paracolias Berger, 1986

Lambillionea 86(7/8, suppl.): 24, 34. Type-species: Colias dimera E. Doubleday, [1847], in: Gen. diurn. Lepid. (1): pl. 9, fig. 3; (7): 74, no. 15, by original designation.

=    Asiocolias Korb, 2005

Cat. Butt. ex USSR: 20. Type-species: Colias christophi Grum-Grshimaïlo, 1885, in: Romanov, Mém. Lépid. 2: 220, by original designation.

Ganura Zetterstedt, [1839]

Ins. Lappon. (5): col. 903. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71. This name was published in synonymy and is unavailable (Code Article 11.6).

 

Colias palaeno (Linnaeus, 1761)

Fauna Svecica (ed. 2): 272, no. 1041.

Original Combination: Papilio [Heliconius] Palaeno

Type Locality: “Pteride rarissime Uppsaliae, frequentior in Finlandia” [Uppsala, South Uppland, Sweden and Finland]

Types: Syntypes in LSUK.

Most of the Palaearctic synonymy is not included.

            =    philomene Hübner, [1805]

Samml. europ. Schmett. 1: pl. 117, figs. 602 ♂ D, 603 ♂ V [1805]; pl. 147, figs. 740 ♀ D, 741 ♀ V [1816].

Original Combination: [Papilio] Philomene

Type Locality: “Lappland” [Norway and Sweden]

Types: Location of type(s) not known, probably lost.

            =    lapponica Staudinger, 1871

In: Staudinger and Wocke, Cat. Lepid. Europ. (2nd ed.): 5, no. 58a; cited Herrich-Schäffer (1848), Syst. Bearb. Schmett. Europa 1(36): pl. 85, figs. 403 ♀ D, 404 ♀ V.

Original Combination: [Colias Palaeno] v. Lapponica

Type Locality: “Lap. Ross. s.” [Murmanskaya Oblast, Russia]

Types: Location of type(s) not known.

 

Colias palaeno chippewa W. H. Edwards, 1870

Syn. Butts. N. Am.: 8, no. 15.

Original Combination: Colias Chippewa

Type Locality: “Great Slave Lake” [Northwest Territories]

Proposed to replace Colias helena W. H. Edwards, 1863, preoccupied, with the same data (Code Article 60.3).

Some authors have treated chippewa as a species distinct from palaeno (e.g., Maey (1986), Mitt. Arbeitsg. westfälischer Lepid. Entomol. Ver. Beiheft 1: 70, Tuzov (1996), Actias 2(1/2): 106). These have been anecdotal and a thorough analysis of taxa occurring on both sides of the Bering Straits is necessary; see Grieshuber and Worthy (2004), Helios 5: 32-36.

                         =    ‡helena W. H. Edwards, 1863

Proc. Entomol. Soc. Philad. 2(2): 78, 80, no. 3; figured by W. H. Edwards (1868), Butts. N. Am. [1](2): pl. Colias 1, figs. 5 ♂ D, 6 ♂ V, 7 ♀ D.

Original Combination: Colias Helena

Type Locality: “Mackenzie's River”; lectotype from “Slave Lake”; defined as “near the west end of Great Slave Lake, North West Territory, Canada” by F. Brown (1973), Trans. Am. Entomol. Soc. 99(1): 54.

Types: Lectotype in CMNH, designated by F. Brown (1973), Trans. Am. Entomol. Soc. 99(1): 55, fig. 11 ♂ [not ♀] D&V; figured by W. Holland (1931), Butt. Book (rev. ed.): pl. 68, fig. 30 ♂ D.

Preoccupied by Colias helena Herrich-Schäffer, 1844, Europ. Schmett. 1(7): pl. 45, figs. 206-207.

                         =    ‡”kohlsaati” (Gunder, 1931)

Bull. South. Calif. Acad. Sci. 30(2): 45.

Original Combination: Eurymus chippewa form ♀ kohlsaati

Type Locality: “Mt. McKinley Nat. Park, Alaska” [Denali National Park]

Types: “Holotype” in AMNH.

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