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Colias tyche kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001
(Booth's Sulphur)

Type Specimens photo collection:


 

Distribution and Larval Foodplants:
 

[olga is preoccupied] Seward Peninsula, AK

Synonymy 

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Colias [Fabricius], 1807

In: Illiger, Mag. f. Insektenk. 6: 284, no. 24. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by designation of I.C.Z.N. Opinion 146.

      =    ‡Eurymus Horsfield, [1829]

Descr. Cat. Lepid. Ins. Mus. E. India Co. (2): 129, 134. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation. Preoccupied by Eurymus Rafinesque, 1815, Analyse Nat.: 117, and a junior objective synonym of Colias.

=    Scalidoneura Butler, 1871

Proc. Zool. Soc. Lond. 1871(1): 250-251. Type-species: Scalidoneura hermina Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 251, pl. 19, fig. 5, by original designation.

=    Eriocolias J. Watson, 1895

Entomol. 28(385): 167. Type-species: Papilio edusa Fabricius, 1787, Mantissa Ins. 2: 23, no. 240, by original designation (= Papilio croceus Geoffroy (1875), in: Fourcroy, Entomol. Paris 2: 250, no. 48).

      =    ‡Coliastes Hemming, 1931

Entomol. 64(12)[823]: 273. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71, by original designation; a junior objective synonym of Colias.

=    Mesocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias vauthierii Guérin-Méneville, [1830], in: Duperrey, Voy. Coquille, 1(Zool.)Atlas(18): pl. Ins. 15, fig. 2, by original designation.

=    Protocolias Petersen, 1963

J. Res. Lepid. 1(2): 144-145. Type-species: Colias imperialis Butler, 1871, Proc. Zool. Soc. Lond. 1871(1): 250, pl. 19, fig. 2, by original designation (= Colias ponteni Wallengren, 1860, Wien. entomol. Monatschr. 4(2): 33, no.1; synonym).

=    Palaeocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21-22, 32. Type-species: Colias ponteni Wallengren, 1860, Wien. Entomol. Montaschr. 4(2): 33, by original designation.

=    Neocolias Berger, 1986

Lambillionea 86(7/8, suppl.): 21, 32. Type-species: Papilio erate Esper, 1805, Die Schmett. 1(suppl.)(2)(11): 13-14, pl. 119 (Cont. 74), fig. 3, by original designation.

=    Eucolias Berger, 1986

Lambillionea 86(7/8, suppl.): 22, 33. Type-species: Papilio palaeno Linnaeus, 1761, Fauna Svecica (ed. 2): 272, no. 1041, by original designation.

=    Similicolias Berger, 1986

Lambillionea 86(7/8, suppl.): 23, 33. Type-species: Papilio lesbia Fabricius, 1775, Syst. Entomol.: 477, no. 149, by original designation.

=    Paracolias Berger, 1986

Lambillionea 86(7/8, suppl.): 24, 34. Type-species: Colias dimera E. Doubleday, [1847], in: Gen. diurn. Lepid. (1): pl. 9, fig. 3; (7): 74, no. 15, by original designation.

=    Asiocolias Korb, 2005

Cat. Butt. ex USSR: 20. Type-species: Colias christophi Grum-Grshimaïlo, 1885, in: Romanov, Mém. Lépid. 2: 220, by original designation.

Ganura Zetterstedt, [1839]

Ins. Lappon. (5): col. 903. Type-species: Papilio hyale Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 469, no. 71. This name was published in synonymy and is unavailable (Code Article 11.6).

 

Colias tyche (Böber, 1812)

Mém. Soc. Imp. Nat. Moscou 3: 21, no. 7, pl. 1, figs. 3 ♂ D, 4 ♂ V; 320 (index, binomial combination).

Original Combination: [Papilio] Dan[aus]. Tyche

Type Locality: “Sibérie” implied from the title of the paper, but stated to be “des environs du lac Baïkal” by Böber, (1809), Mém. Soc. Imp. Nat. Moscou 2: 303. [Republic of Buryatia, Russia]

Types: Syntypes probably lost.

Grieshuber (1998), Lambillionea 98(1)(Tome 2): 126-134, 98(3)(Tome 2): 453-468, and (1998) 98(4)(Tome 1): 509-522, revised this species, and proposed the following synonymy.

            =    melinos Eversmann, 1847

Bull. Soc. Imp. Nat. Moscou 20(2)(3): 72-73, no. 7, pl. 3, figs. 3 ♂ D, 4 ♂ V, 5 ♀ D, 6 ♀ V.

Original Combination: Colias Melinos

Type Locality: “Sibiria orientali”; suggested to be “environs of Ulan-Ude” by Grieshuber (2004), Helios 5: 8. [Republic of Buryatia, Russia]

Types: Lectotype in ZIN, designated by Grieshuber (2004), Helios 5: 6-7, pl. 1, fig. 12 ♂ D.

            =    chloe Eversmann, 1847

Bull. Soc. Imp. Nat. Moscou 20(2)(3): 73-74, no. 8, pl. 4, figs. 1 ♂ D, 2 ♂ V.

Original Combination: Colias Chloë

Type Locality: “Sibiria orientali, in provincia Irkutskensi, circa Kiachtam”; suggested to be “environs of Ulan-Ude” by Grieshuber (2004), Helios 5: 6. [Republic of Buryatia, Russia]

Types: Lectotype in ZIN, designated by Grieshuber (2004), Helios 5: 5-6, pl. 1, fig. 10 ♂ D.

            =    magna Rühl, 1893

In: Heyne, Palae. Grossschmett. 1(3): 153.

Original Combination: [Colias Melinos] var. magna

Type Locality: “Baikalien” [Republic of Buryatia, Russia]

Types: Location type(s) not known.

            =    alpina Elwes, 1899

Trans. Entomol. Soc. Lond.(4)1899(3): 304, 319-320.

Original Combination: Colias melinos var. alpina

Type Locality: “the high bare mountains south of the Tchuja Steppe” Elwes' earlier itinerary (pages 298-299) allows the further definition to "Tachety River, Tschuja Steppe, 30 mi S Kuch Agatch, Altai Republic, Altay Kray, Russia." [Altai Mountains, Altai Republic, Altay kray, Russia]

Types: Syntypes in BMNH, ZMHB, and ZIN.

            =    vitimensis Austaut, 1899

Naturaliste (2)13(307): 284.

Original Combination: Colias Melinos V[ar]. Vitimensis

Type Locality: “des règions alpines de la Siberie orientale, où la rivière Vitim va se jeter dans le fleuve Lèna” [Sakha Autonomous Republic, Russia]

Types: Syntypes possibly in BMNH.

            =    herzi Staudinger, 1901

In: Staudinger and Rebel, Cat. Lepid. Palae. (3rd ed.): 15, no. 90a; cited Colias melinos Herz (1899), Dtsche. Entomol. Z. “Iris” 11(2): 235.

Original Combination: [Colias Melinos] v. Herzi

Type Locality: “Sib. S. or. (Vilui)”; lectotype from “Podeibi” [Bodajbo, Irkutskaya oblast, Russia]

Types: Lectotype in ZMHB, designated by Grieshuber, 2006, Helios 7: 79, pl. 16, fig. 4 ♂ D.

            =    deckerti Verity, [1909]

Rhop. palae. (21-23): 236 [30 Apr 1909]; (30-36): pl. 72, 27 ♂ D, 28 ♀ D; ind. syst.: xxxvi [31 Oct 1911].

Original Combination: [Colias melinos] var. deckerti

Type Locality: “Monts Apfel”; suggested to be “Pokrovka” by Grieshuber (1998), Lambillionea 98(1)(Tome 2): 130-131, no. 2. [Pokrofka, Chita Oblast, Russia]

Types: Syntypes in BMNH and ZMKU; figured by Chikolovets et al. (2002), Butts. Transbaikal Siberia: pl. 7, figs. 8 ♂ D, 23 ♀ V.

                   =    ‡”chryseis” Verity, [1911]

Rhop. palae. (30-36): 354, pl. 70, figs. 33 ♂ D, 34 ♀ D; ind. syst.: xxxvi.

Original Combination: [Colias melinos] race chryseis

Type Locality: “l’Amour [Amur Oblast, Russia]

Types: Syntypes in BMNH.

                   =    ‡”montana” Verity, [1911]

Rhop. palae. (30-36): 354, pl. 70, figs. 37 ♂ D, 38 ♀ D; ind. syst.: xxxvi.

Original Combination: [Colias melinos] race montana

Type Locality: “l’Altaï” implied from the text, “Montagnes de Tchuja, 1800-2400 m., S.E. l’Altaï” from the legend to plate 70. [Altai Republic, Altay Kray, Russia]

Types: Syntypes in BMNH.

            =    montana Bollow, 1930

In: Seitz (1931), Grossschmett. Erde 1(Suppl.)(14): 108.

Original Combination: [Colias melinos chryseis

Bollow adopted Colias melinos race montana Verity [1911] to represent a subspecies-level taxon, with the same data (Code Articles 10.2, 45.5.1).

            =    chryseis Bollow, 1930

In: Seitz (1931), Grossschmett. Erde 1(Suppl.)(14): 108-109.

Original Combination: [Colias melinos chryseis

Bollow adopted Colias melinos race chryseis Verity [1911] to represent a subspecies-level taxon, with the same data (Code Articles 10.2, 45.5.1).

            =    ludmilla Hemming, 1933

Entomol. 66(12)[847]: 278, no. 23.

Original Combination: Colias melinos ludmilla

Proposed to replace Colias melinos race chryseis Verity, 1911, with the same data (Code Article 60.3).

            =    jeholensis Matsumura, 1939

Bull. Biogeogr. Soc. Japan 9(20): 344, no. 19; 348, no. 19, pl. 13, fig. 16 ♂ D (actually fig. 6).

Original Combination: Colias herzi form jeholensis

Type Locality: “Jehol...Mt. Kingan” [Nei Mongol Autonomous Region, China]

Types: Syntypes probably in EIHU.

            =    relicta Kurentsov, 1970

Butts. Far East USSR: 38, fig. 28(4) ♂ D, fig. 29(8) ♂ genitalia.

Original Combination: C[olias]. m[elinos]. relicta

Type Locality: [the mountains at the source of the Okhota River, where it favors a few peaks in the Verkhoyansk Range, to an elevation of 3000 meters above sea level]; holotype from [Burgakhchan]. [Magadan Oblast, Chukotka Autonomous Okrug, Russia]

Types: Holotype in the IPBV; registered by Azarova (1986), Syst. Ecol. Lepid. Far East USSR: 122, no. 4; figured by Churkin et al. (2001), Helios 2: pl. 8, figs. 13 ♂ D, 16 ♂ V.

 

Colias tyche kolosovae Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001

Atalanta 32(1/2):262.

Original Combination: Colias tyche kolosovae

Proposed as a replacement name for Colias tyche olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001, preoccupied, with the same data (Code Article 60.3).

                         =    ‡olga Churkin, Grieshuber, Bogdanov & Zamolodchikov, 2001

Helios 2: 109-110, pl. 9, figs. 1 ♂ D, 2 ♂ D (holotype); 3 ♂ D, 4 ♂ D, 5 ♀ D, 15 ♂ V.

Original Combination: Colias tyche olga

Type Locality: “Russia, East Chukotka, 20 km SE Ioni Lake, Gilmimlviem, [65o 48' N, 173o 22' W]” [Magadan Oblast, Chukotka Autonomous Okrug, Russia]

Types: Holotype in DSM.

Preoccupied by Colias olga Romanoff, 1882, Romanoff, 1882, Horae Soc. Entomol. Ross. 17(3/4): 127, pl. 4, figs. 1-3.

Bibliography

 

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