Phoebis sennae marcellina (Cramer, 1777)

Phoebis sennae marcellina (Cramer, 1777)
(Cloudless Sulphur)

Distribution and Larval Foodplants:

SW USA (strays far N), S TX through Panama, most S. America

(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Phoebis Hübner, [1819]

Verz. bekannt. Schmett. (7): 98. Type-species: Phoebis cypris Hübner, [1819], Verz. bekannt. Schmett. (7): 98, no. 1044 (and unjustified emendation of Papilio cipris Cramer, 1777, Uitl. Kapellen 2(9): 5, pl. 99, figs. E, F; (16): 148 (index); = Papilio argante Fabricius, 1775, Syst. Entomol.: 470, no. 116; synonym), by designation of Butler (1873), Lepid. Exot. (13): 155.

= ‡Colias Hübner, [1819]

Verz. bekannt. Schmett. (7): 99. Type-species: Papilio sennae Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 470, no. 72, by designation of Hemming (1939), Proc. R. Entomol. Soc. Lond. (B)8(7): 136. Preoccupied by Colias [Fabricius], 1807, in: Illiger, Mag. f. Insektenk. 6: 284.

= Callidryas Boisduval & Le Conte, [1830]

Hist. Lépid. Amér. sept. (8): 73. Type-species: Papilio eubule Linnaeus, 1767, Syst. Nat. (ed. 12) 1(2): 764, no. 102 (= Papilio sennae Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 470, no. 72; synonym), by designation of Blanchard (1840), in: Castelnau, Hist. Nat. Ins. 3: 432.

= ‡Metura Butler, 1873

Lepid. Exot. (13): 155. Type-species: Callidryas bracteolata Butler, 1865, Proc. Zool. Soc. Lond. 1865(2): 458, pl. 26, fig. 6 (= Colias neocypris Hübner, [1823], Samml. exot. Schmett. 2: pl. [136], figs. 1, 2; synonym), by designation of Scudder (1875), Proc. Am. Acad. Arts Sci., Boston 10(2): 218, no. 695. Preoccupied by Metura Walker, 1855, in: Allen's Nat. Libr., Lepid. 2: 229.

= Parura W. F. Kirby, 1896

In: Allen, Nat. Libr. Lepid. 2: 229. Type-species: Callidryas bracteolata Butler, 1865, Proc. Zool. Soc. Lond. 1865(2): 458, pl. 26, fig. 6 (= Colias neocypris Hübner, [1823], Samml. exot. Schmett. 2: pl. [136], figs. 1, 2; synonym), as a replacement name. Proposed to replace Metura Butler, 1873, preoccupied (Code Articles 60.3, 67.8).

Phoebis sennae (Linnaeus, 1758)

Syst. Nat. (ed. 10) 1: 470, no. 72; cited Sloane (1725), Voy. Mad. Barb. Niev. St. Christ. Jam. 2: pl. 236, figs. 11, 12, and Merian (1705), Ins. Surinam.: 58, pl. 58, among others.

Original Combination: P[apilio]. D[anaus Candidus]. Sennae

Type Locality: “in Cassia Americes”; possibly “Jamaica” from reference to Sloane.

Types: Type(s) apparently lost, as noted by Honey and Scoble (2001), Zool. J. Linn. Soc. 132(3): 379.

= eubule (Linnaeus, 1767)

Syst. Nat. (ed. 12) 1(2): 764, no. 102; cited G. Edwards (1760) Glean. Nat. Hist. 2: pl. 304.

Original Combination: P[apilio]. D[anaus Candidus] Eubule

Type Locality: “Carolina”; suggested to be “vic. of Charleston, South Carolina” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 86. [Charleston County]

Types: Type(s) apparently lost.

Lamas (2004d), Atl. Neotrop. Lepid. Chcklst. Part 4A: 104, determined this synonymy.

= drya (Fabricius, 1775)

Syst. Entomol.: 478, no. 153.

Original Combination: P[apilio]. D[anaus]. C[andidus]. Drya

Type Locality: “America”; suggested to be “vic. of Charleston, South Carolina” by L. Miller and F. Brown (1981), Mem. Lepid. Soc. (2): 86. [Charleston County]

Types: Type(s) probably lost.

= ‡”pallida” (Cockerell, 1889)

Entomol. 22(308): 5; cited French (1886), Butts. E. U. S.: 122.

Original Combination: Callidryas sennae pallida ♀

Type Locality: Not stated.

Types: “Type” is apparently lost.

= ‡”lichas” (d'Almeida, 1922)

Mélanges Lépid. 1: 56, no. 43; cited Boisduval and Le Conte [1830], Hist. Lépid. Amér. sept. (8): pl. 24, fig. 3 ♀ V.

Original Combination: [Catopsilia Eubule] Form. aberr. Lichas

Type Locality: Not specifically stated, but probably “Georgia” based on the figures by John Abbot in Boisduval and Le Conte.

Types: Described, at least in part, from a John Abbot drawing. The specimens upon which this drawing was based are presumed lost. A “syntype” is in USNM, but Calhoun (2004), J. Lepid. Soc. 58(3): 156, discussed the dubious nature of such “types” from the Boisduval collection.

= ‡”sennalba” F. Brown, 1929

Am. Mus. Novit. (368): 8-9.

Original Combination: Phoebis eubule sennae female form sennalba

Type Locality: “7 kilometers north of Viñales, Cuba” [Pinar del Rio]

Types: “Holotype” in AMNH.

= ‡”browni” W. D. Field, 1936

J. Entomol. & Zool. 28(2): 20.

Original Combination: Phoebis sennae race eubule female f. browni

Type Locality: “Lawrence, Kansas” [Douglas County]

Types: “Holotype” in USNM.

‡”alba” Scott, 1986

Butts. N. Am.: 204, no. 50.

Original Combination: [Phoebis sennae ♀] form alba

Proposed as a “form” with no nomenclatural standing; a nomen nudum.

‡”rosa” Scott, 1986

Butts. N. Am.: 204, no. 50.

Original Combination: [Phoebis sennae] winter form rosa

Proposed as a “form” with no nomenclatural standing; a nomen nudum.

Phoebis sennae marcellina (Cramer, 1777)

Uitl. Kapellen 2(14): 103, pl. 163, fig. C (A ♀ D, B ♀ V, C ♂ V); (16): 149 (index).

Original Combination: Pap[ilio]. Dan[aus]. Cand[idus]. Marcellina

Type Locality: “Surinamen” “Suriname”

Types: Syntypes (#665067, 665068) in BMNH.

= hyperici (Sepp, [1832])

Surinaam. Vlinders [1](5): 45, pl. 19.

Original Combination: Papilio Hyperici

Type Locality: “Surinam” implied from the title of the work.

Types: Type(s) lost.

= rhadia (Boisduval, 1836)

Spec. gén. Lépid. 1: 617, no. 11.

Original Combination: Callidryas Rhadia

Type Locality: “Senegal, île Maurice” in error. [Mauritius, West Africa]

Types: Syntypes in BMNH.

= yamana (Reakirt, [1864])

Proc. Entomol. Soc. Philad. 2(3): 352, no. 3.

Original Combination: C[allidryas]. Yamana

Type Locality: “Honduras”

Types: Type(s) may be in CMNH or lost.

= ‡”schausi” (Oberthür, 1912)

Étud. Lépid. Comp. 6: 328, pl. 126, fig. 1116 ♂ D&V (“holotype”).

Original Combination: Callidryas Eubule-Schausi

Type Locality: “Nouvelle- Fribourg” [Nova Friburgo, Rio de Janeiro, Brazil]

Types: “Holotype” in BMNH.

The text clearly indicates that this taxon was described as an individual variant.

= ‡”fugax” (d'Almeida, 1922)

Mélanges Lépid. 1: 57-58, no. 44; figured by d'Almeida (1940), Arq. Zool. São Paulo 1(3): pl. 4, fig. 7.

Original Combination: [Catopsilia Eubule] form. Fugax

Type Locality: Not stated; “neotype” from “Piedade, rio de Janeiro” [Brazil]

Types: “Neotype” (#1089) in UFPC, designated by Mielke and Casagrande (1985), Revta. bras. Entomol. 29(2): 323, fig. 2 ♂ D&V, but this designation was not in accordance with Code Articles 75.2 and 75.3.

The text clearly indicates that this taxon was described as an individual variant.

= ‡”giacomellii” Köhler, 1923

Z. wiss. InsektBiol. 18(12)(Sonderheft): 16, unnumbered fig. ♀ V.

Original Combination: Catopsilia eubule f. Giacomellii

Type Locality: “La Rioja” [Argentina]

Types: “Syntype” in MLP.

= ‡”minor” Dufrane, 1947

Bull. Annls. Soc. Entomol. Belg. 83(1/2): 64.

Original Combination: [Phoebis sennae marcellina] minor

Type Locality: Not stated; “lectotype” from “Bolivia.”

Types: “Lectotype” in IRSN, designated by Lamas (1981), Revta. Cienc. (Lima) 73(1): 58.

= ‡”albescens” Dufrane, 1947

Bull. Annls. Soc. Entomol. Belg. 83(1/2): 64.

Original Combination: [Phoebis sennae marcellina] ab. albescens

Type Locality: “Brésil”

Types: “Holotype” in IRSN.

= ‡”major” Dufrane, 1947

Bull. Annls. Soc. Entomol. Belg. 83(1/2): 65.

Original Combination: [Phoebis sennae marcellina] major

Type Locality: “Brésil”; lectotype from “Peru.”

Types: “Lectotype” in IRSN, designated by Lamas (1981), Revta. Cienc. (Lima) 73(1): 56.

Bibliography

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