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Phyciodes c. cocyta (Cramer, 1777)
(Northern Crescent)

Pinned Specimens
photo collection:

Live Adults
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Larval Foodplants and Habitats
photo collection:

Distribution and Larval Foodplants:

Asteraceae: Symphyotrichum laeve var. laeve (L.) A.& D. Löve


(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Phyciodes Hübner, [1819]

Verz. bekannt. Schmett. (2): 29. Type-species: Papilio cocyta Cramer, 1777, Uitl. Kapellen 2(9): 7, pl. 101, figs. A, B; (16): 148 (index), by designation of Scudder (1872), 4th Ann. Rep. Peabody Acad. Sci. (1871): 46. The arrangement of species in this genus follows Wahlberg et al. (2003), Syst. Entomol. 28(2): 257-273.


Phyciodes cocyta (Cramer, 1777)


Phyciodes cocyta cocyta (Cramer, 1777)

Uitl. Kapellen 2(9): 7, pl. 101, figs. A ♂ D, B ♂ V (fig. C ♀ V is not the same species); (16): 148 (index).

Original Combination: Pap[ilio]. Nymph[alis]. Phaler[ata]. Cocyta

Type Locality: Not stated for figs. A and B, “Surinamen” “Suriname” for fig. C; neotype from “Black Rock, Cape Breton, Nova Scotia.”

Types: Neotype in AMNH, designated by Scott (1994), Papilio (n.s.) 7: 36, fig. 111 ♂ D.

                         =    ‡morpheus (Fabricius, 1775)

Syst. Entomol.: 530, no. 370.

Original Combination: P[apilio]. P[lebejus]. R[uralis]. Morpheus

Type Locality: “America boreali”

Types: Type(s) probably lost.

Preoccupied by Papilio morpheus Pallas, 1771, Reise verschied. Prov. Russisich. Reichs 1, Anhang: 471, no. 64. Fabricius (1793), Entomol. Syst. 3(1): 155-156, no. 479, cited this name in association with Papilio cocyta Cramer, 1777.

                   =    arctica dos Passos, 1935

Can. Entomol. 67(4): 87.

Original Combination: Phyciodes tharos arctica

Type Locality: “Port au Port, Table Mountain, Newfoundland” [Newfoundland and Labrador]

Types: Holotype in AMNH.


Cramer, Pieter. 1775-1790. Uitlandische Kapellen, voorkomende in de drei Waereld-Deelen Asia, Africa en America, by een verzameld en bescreeven. [Dutch Title]. Papilions exotiques des trois parties du monde l"Asie, l'Afrique et l'Amerique. [French Title]. Amsterdam, Baalde,and Utrecht, Wild. 4 vols xxx + 16 + 765 pp.; 400 pl. {1775-1790} [description of Papilio cocyta

dos Passos, Cyril Franklin. 1935. Some butterflies of southern Newfoundland with descriptions of new subspecies (Lepid. Rhopal.). Canadian Entomologist 67(4): 82-88. {4 May 1935} [description of Phyciodes tharos arctica

Fabricius, Johann Christian. 1775. Systema Entomologiae, Sistens Insectorvm Classes, Ordines, Genera, Species, Adjectis Synonymis, Locis, Descriptionibvs, Observationibvs. Flensburgh and Leipzig; Library Kortii, 8vo: [iv] + [xii] + xvi] + 832 pp. (Appendix: 816-832) {[17 Apr] 1775} [description of Papilio morpheus

Forbes, William Trowbridge Merrifield. 1945. The genus Phyciodes (Lepidoptera, Nymphalinae). Entomologica Americana (n.s.)24: 139-207. {Oct 1944, 24 May 1945} 

Hall, Arthur. 1928-1930. A revision of the genus Phyciodes Hubn. (Lepidoptera, Nymphalidae). Bulletin of the Hill Museum 2(2)(supplement 1): 1-20, pls. 1-3 {18 Jun 1928} 2(3)(supplement): 21-44 {17 Nov 1928}; 3(1)(supplement): 45-94 {30 Apr 1929}; 3(2)(supplement): 95-146 {29 Aug 1929}; 3(3)(supplement): 147-170 {30 Dec 1929}; 4(1)(supplement): 171-204 {28 Jul 1930}; 4(2)(supplement): 205-206; add. Page {29 Dec 1930} 

Higgins, Lionel George. 1981. A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae (Lepidoptera: Nymphalidae). Bulletin of the British Museum (Natural History) (Entomology) 43(3): 77-243, 490 figs. {29 Oct 1981} 

Layberry, Ross A., Peter W. Hall, J. Donald Lafontaine (plates by John T. Fowler). 1998. The butterflies of Canada. Toronto, Buffalo & London; University of Toronto Press: 280 pp., 32 pls., 294 maps and illus. {1998} 

Scott, James A. 1994. Biology and systematics of Phyciodes (Phyciodes). Papilio (n.s.) 7: 1-120, 227 figs., 12 tables {18 Nov 1994} 

Scott, James A. 1998. Phyciodes (Phyciodes): New discoveries, new subspecies, and convergence. Papilio (New Series) 10: 1-42, 41 figs., 5 tbls. {20 Feb 1998} 

Scott, James A. 2006. Phyciodes (Phyciodes): more progress. Papilio (New Series) 13: 38 pp., 5 tbls., cartoon {28 Apr 2006}

Wahlberg, Niklas, Rita Oliviera and James A. Scott. 2003. Phylogenetic relationships of Phyciodes butterfly species (Lepidoptera: Nymphalidae): complex mtDNA variations and species delimitations. Systematic Entomology 28(2): 257-273, 4 figs., 2 tbls., 1 app. {Apr 2003}

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