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Phyciodes pallida barnesi Skinner, 1897
(Pale Crescent)

Pinned Specimens photo collection Page 1:


Pinned Specimens photo collection Page 2:

Distribution and Larval Foodplants:

Asteraceae: Cirsium arizonicum (Gray) Petrak, Cirsium neomexicanum Gray, Cirsium undulatum (Nutt.) Spreng.


(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Phyciodes Hübner, [1819]

Verz. bekannt. Schmett. (2): 29. Type-species: Papilio cocyta Cramer, 1777, Uitl. Kapellen 2(9): 7, pl. 101, figs. A, B; (16): 148 (index), by designation of Scudder (1872), 4th Ann. Rep. Peabody Acad. Sci. (1871): 46. The arrangement of species in this genus follows Wahlberg et al. (2003), Syst. Entomol. 28(2): 257-273.


Phyciodes pallida (W. H. Edwards, 1864)

Proc. Entomol. Soc. Philad. 2(4): 501, 505, no. 6.

Original Combination: Melitaea Pallida

Type Locality: “Texas. Kansas”; neotype from “Flagstaff Mt., Boulder Colo.” [Boulder County, Colorado]

Types: Neotype in CMNH, designated by F. Brown (1966), Trans. Am. Entomol. Soc. 92(3): 448, fig. 28 ♀ D&V.

Species-genus gender conformation is pallidus.

=    mata (Reakirt, 1866)

Proc. Entomol. Soc. Philad. 6(1): 142.

Original Combination: Eresia mata

Type Locality: “Rocky Mountains, Colorado Terr.”; defined as “Flagstaff Mt., Boulder Colo.” by Scott (1994), Papilio (n.s.) 7: 11. [Boulder County]

Types: Holotype in FMNH; figured by Strecker (1874), Lepid. Rhop. Het. (8): pl. 8, fig. 11 ♀ D&V.


Phyciodes pallida barnesi Skinner, 1897

Can. Entomol. 29(6): 155.

Original Combination: Phyciodes Barnesi

Type Locality: “Glenwood Springs, Colo.” [Garfield County, Colorado]

Types: Syntypes (#7037) in ANSP, one figured by W. Holland (1898), Butt. Book: 156, pl. 18, fig. 5 ♂ D.


Barnes, William and James Halliday McDunnough. 1916. Notes on North American diurnal Lepidoptera. Contributions to the Natural History of the Lepidoptera of North America 3(2): 49-156, 8 pls. {15 Dec 1916} 

Ferris, Clifford Duras and Frederick Martin Brown. 1981. Butterflies of the Rocky Mountain States. Oklahoma; Univ. Oklahoma Press: 442 pp., 4 cpls., many figures [contributions from Donald Eff, Scott Ellis, Michael Fisher, Lee D. Miller, James Scott and Raymond Stanford] {1981} 

Forbes, William Trowbridge Merrifield. 1945. The genus Phyciodes (Lepidoptera, Nymphalinae). Entomologica Americana (n.s.)24: 139-207. {Oct 1944, 24 May 1945} 

Gillham, Nicholas W. and Paul Ralph Ehrlich. 1954. The butterfly types of Henry Skinner and co-authors in the Academy of Natural Sciences of Philadelphia. Transactions of the American Entomological Society 80(3/4): 91-117 {20 Aug 1954} 

Guppy, Crispin S. & Jon H. Shepard. 2001. Butterflies of British Columbia. Including western Alberta, southern Yukon, the Alaska panhandle, Washington, northern Oregon, northern Idaho, northwestern Montana. Vancouver-Toronto; Royal British Columbia Museum, Univ, British Columbia Press: 414, 75 figs., text figures, maps. {2001} 

Hall, Arthur. 1928-1930. A revision of the genus Phyciodes Hubn. (Lepidoptera, Nymphalidae). Bulletin of the Hill Museum 2(2)(supplement 1): 1-20, pls. 1-3 {18 Jun 1928} 2(3)(supplement): 21-44 {17 Nov 1928}; 3(1)(supplement): 45-94 {30 Apr 1929}; 3(2)(supplement): 95-146 {29 Aug 1929}; 3(3)(supplement): 147-170 {30 Dec 1929}; 4(1)(supplement): 171-204 {28 Jul 1930}; 4(2)(supplement): 205-206; add. Page {29 Dec 1930} 

Higgins, Lionel George. 1981. A revision of Phyciodes Hübner and related genera, with a review of the classification of the Melitaeinae (Lepidoptera: Nymphalidae). Bulletin of the British Museum (Natural History) (Entomology) 43(3): 77-243, 490 figs. {29 Oct 1981} 

Holland, William Jacob. 1898. The butterfly book; a popular guide to a knowledge of the butterflies of North America. Garden City, New York, Doubleday & McClure Co.  xx + 382 pp., 48 pls., 183 + [1] figs. {1898} 

Holland, William Jacob. 1931. The butterfly book. New and thoroughly revised edition.  A popular and scientific manual, describing and depicting all the butterflies of the United States and Canada. Garden City, New York; Doubleday, Doran & Co.  xii + 424 pp., 77 pls., 198 + [1] text figures. {1931}

Layberry, Ross A., Peter W. Hall, J. Donald Lafontaine (plates by John T. Fowler). 1998. The butterflies of Canada. Toronto, Buffalo & London; University of Toronto Press: 280 pp., 32 pls., 294 maps and illus. {1998} 

Pyle, Robert Michael. 2002. Butterflies of Cascadia. Seattle; Seattle Audubon Society: 420 pp., figs., drawings, photos. {2002} 

Röber, Johannes Karl Max 1913-1914. Gattung Phyciodes. In: A. Seitz (Ed.), Die Gross-Schmetterlinge der Erde 5(162): pl. 92 {5 Jun 1913}; (183): 434-440, pl. 90 {10 Dec 1913}; (188): 449-450, pls. 88, 89. {[before 10 Feb] 1914}; (189): pl. 91. {14 Apr 1914} 

Scott, James A. 1994. Biology and systematics of Phyciodes (Phyciodes). Papilio (n.s.) 7: 1-120, 227 figs., 12 tables {18 Nov 1994} 

Scott, James A. 1998. Phyciodes (Phyciodes): New discoveries, new subspecies, and convergence. Papilio (New Series) 10: 1-42, 41 figs., 5 tbls. {20 Feb 1998} 

Scott, James A. 2006. Butterfly hostplant records, 1992-2005, with a treatise on the evolution of Erynnis, and a note on new terminology for mate-locating behavior. Papilio (New Series) 14: 74 pp., cartoon {28 Apr 2006} 

Skinner, Henry. 1897. Notes on Rhopalocera, with descriptions of new species and varieties. Canadian Entomologist 29(6): 154-156. {June 1897} [description of Phyciodes Barnesi

Tilden, James William. [1970]. Concerning the names and status of certain North American members of the genus Phyciodes. Journal of Research on the Lepidoptera 8(3): 94-98. {3 Sep 1969 [21 Dec 1970]}

Wahlberg, Niklas, Rita Oliviera and James A. Scott. 2003. Phylogenetic relationships of Phyciodes butterfly species (Lepidoptera: Nymphalidae): complex mtDNA variations and species delimittions. Systematic Entomology 28(2): 257-273, 4 figs., 2 tbls., 1 app. {Apr 2003} 

Warren, Andrew D. 2005. Butterflies of Oregon: their taxonomy, distribution, and biology. Fort Collins; Contributions of the C.P. Gillette Museum of Arthropod Diversity, Colorado State University: 405 pp., 3 maps. {15 Mar 2005}

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