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Plebejus shasta minnehaha (Scudder, 1874)
(Shasta Blue)

Type Specimens photo collection:

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(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Plebejus Kluk, 1780

Zwierz. Hist. Nat. Pocz. Gospod. 4: 89-92, no. 148. Type-species: Papilio argus Linnaeus, 1758, Syst. Nat. (ed. 10) 1: 483, no. 152, by designation of Hemming (1933), Entomol. 66(845): 224. I.C.Z.N. Opinion 278 placed Plebejus on the Official List of Generic Names in Zoology as name no. 705. This name has been spelled various ways; the spelling given in the Official List is the proper spelling by rule (Code Article 79.4.1). Bálint et al. (2001), Folia Entomol. Hung. 62: 177-184, discussed this issue.


Plebejus shasta (W. H. Edwards, 1862)

Proc. Acad. Nat. Sci. Philad. 14(5): 221, 224-225, no. 8.

Original Combination: Lycaena Shasta

Type Locality: “California”; neotype from “Sheppard Pass Trail West/head of Tyndall Ck. 11-12,200'/, Tulare Co., Calif.”

Types: Neotype in CMNH, designated by F. Brown (1970), Trans. Am. Entomol. Soc. 96(3): 404, fig. 20 ♂ D&V.

J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 27, discussed the neotype designated by F. Brown (1970). They considered it invalid because Behr did not have access to the area from where the neotype was taken, and W. H. Edwards’ description fits the syntypes of nivium and zelmira more closely than that of the neotype. Austin et al. (1998), Syst. W. N. Am. Butts. (7): 115-116, designated a “new” neotype. This is the lectotype of zelmira (see below). This issue needs further examination.

=    ‡ zelmira (C. Felder & R. Felder, 1865)

Reise Novara (2): 282, no. 358, pl. 35, figs. 17 ♂ D, 18 ♂ V, 19 ♀ D.

Original Combination: Lycaena Zelmira

Type Locality: “California, Utah”; defined as “slope south above Donner Pass, 7,500' elevation, Placer County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 90.

Types: Lectotype in BMNH, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (5): 90, figs. 28 ♂ D, 29 ♂ V. This is the “neotype” of shasta, thereby making zelmira a junior objective synonym of shasta (see above).

=    nivium (Boisduval, 1869)

Annls. Soc. Entomol. Belg. 12: 47, no. 25; figured by Oberthür (1913), Étud. Lépid. Comp. 9(1): pl. 238, figs. 1963 ♂ D&V, 1964 ♀ D&V.

Original Combination: Lycaena nivium

Type Locality: “l'Orégon”; defined as “slope south above Donner Pass, 7,500' elevation, Placer County, California” by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 27.

Types: Lectotype in USNM, designated by J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 27, figs. 195 ♂ D, 196 ♂ V.

=    comstocki C. Fox, 1924

Entomol. News 35(4): 140-141.

Original Combination: Plebejus shasta comstocki new variety

Type Locality: “Glacier Point, Yosemite National Park, California” [Mariposa County]

Types: Holotype in AMNH; figured by J. A. Comstock (1927), Butts. Calif.: pl. 54, figs. 27 ♂ D.


Plebejus shasta minnehaha (Scudder, 1874)

Proc. Boston Soc. Nat. Hist. 17(1): 88-89, no. 12.

Original Combination: Agriades minnehaha

Type Locality: “Heart River Crossing, Dakotah Terr., about fifty miles west of the Missouri River”; suggested to be “most probably in Grant Co., North Dakota where the present State Road 49 (between Glen Ullin and Elgin) crosses the Heart River...elevation...1800'” by Ferris (1970), Entomol. News 81(8): 204.

Types: Syntypes (#15977) in MCZ; one figured by Ferris (1976), Bull. Allyn Mus. (36): fig. 16 ♂ D&V.

=    browni Ferris, 1970

Entomol. News 81(8): 204-207, figs. 1, middle row ♂ genitalia, 2, top row left ♂ D (holotype), right ♀ D, bottom row left ♂ V, right ♀ V.

Original Combination: Plebejus shasta browni

Type Locality: “Powder River Pass, 9700', Johnson Co., Wyoming”

Types: Holotype in LACM.



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