Porphyrogenes speciosus Austin and Mielke, new species
(Fig. 101, 102, 133, 153)
Description. Male (Fig. 101, 102) - forewing length = 29.6 mm (holotype); forewing with costal fold,
apex slightly produced, slightly pointed, termen convex apically, then nearly straight, anal margin broadly
convex in middle; hindwing termen broadly convex, tornus produced to very short lobe, vein Rs arising
nearer to end of discal cell than to its base and distad of CuA2 (Fig. 133); dorsum brown, unmarked;
forewing overscaled with orange-brown, heaviest basad in CuA2 and anal cell, anal margin with orange-
brown tuft proximad; hindwing overscaled with orange-brown on basal half posterior to vein Rs; shining
gray speculum in anterior discal cell, proximal 1/4 of Sc+R1-Rs, and proximal 1/2 of costal cell; moderately
long pale yellow-tan recumbent tuft near base of Sc+R1-Rs covering speculum in that cell, semierect pale
yellow-tan tuft arising from near base of discal cell covering speculum in discal cell (Fig. 133); conspicu-
ous erect pale yellow-tan tuft along anterior edge of vein 2A, recumbent orange-brown tuft from posterior
edge of 2A; fringes on both wings brown.
Venter brown, hindwing with slight purplish tinge especially in CuA2-2A; forewing vein 2A curved,
not swollen, surrounded (broadest on cephalic edge) 1/3 distance from base by oval area of modified hair-
like scales (appearing as a brand without magnification, Fig. 133); shining gray-tan speculum in proxi-
mal 1/3 of posterior 2/3 of CuA2-2A and in nearly entire anal cell; hindwing with no indication of discal
macules; cell 2A-3A with deep groove just caudad of vein 2A.
Dorsal head gray-brown with a few greenish scales, palpi gray, eyes red, antennae black, yellow on
venter most extensive distad on shaft, nudum dark red-brown, 28 (n = 1)
segments, dorsal thorax orange-
brown, ventral thorax mixed brown and blue-green, pectus brown with a few greenish scales cephalad,
legs brown, dorsal abdomen dark brown, ventral greased but appearing dark gray.
Genitalia (Fig. 153) - tegumen narrow in lateral view, broad in dorsal view, long, thin and dorso-
caudaloriented process from each side of caudal end, tuft double and sparse; uncus decurved in lateral
view, broadly divided in dorsal view, arms widely spaced, long and parallel, ventral process of uncus more
or less triangular; gnathos shorter than uncus, terminal ends lobate inwardly in ventral view; combined
ventral arms from tegumen and dorsal arms from saccus slightly curved; saccus broad, oriented dorso-
cephalad; valva with costa-ampulla broadly quadrate, harpe long, narrow, bent abruptly dorsad at about
its middle, broadened slightly beyond bend, then narrowing to weakly spiculose caudal end; aedeagus
about length of valva, broad with blunt caudal end; cornuti as short and nearly straight spikes.
Female - unknown.
Type. Holotype male with the following labels: white, handprinted - / Buen Amigo / 27 Apr 95 / MAU
– II Hr. /; white, handprinted - / 297 ? /; white, printed and handprinted - / Genitalic Vial / GTA - 13985
/; red, printed - / HOLOTYPE / Porphyrogenes speciosus / Austin & Mielke /. Deposited for now in the
Type locality. COSTA RICA: Puntarenas; San Luis Monteverde, Buen Amigo.
Etymology. The name means “handsome or imposing” and refers to this species’ large size and
Distribution and phenology. The species is known only from the holotype.
Diagnosis and discussion. Porphyrogenes speciosus is one of the larger species of the genus. The
valva, especially the long and upcurved harpe, of P. speciosus is similar to this structure of P. stresa, P.
zohra, and P. sparus, but the costa-ampulla is broader and more quadrate than on any of those species. In
addition, the uncus of P. speciosus is longer than on those species. The valva of P. sula as illustrated by
Williams and Bell (1940) is somewhat quadrate as on P. speciosus and the harpe appears similarly
expanded near its bend. Porphyrogenes sula, however, is smaller, has different venation and wing shape,
and has blackish tufts anteriorly on the dorsal hindwing;