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Family HESPERIIDAE Latreille, 1809   —   SKIPPERS
Subfamily Eudaminae Mabille, 1877   —   Spreadwing Skippers

Phocides polybius lilea

(Reakirt, [1867])

Guava Skipper

S Texas to Costa Rica

Phocides polybius polybius

(Fabricius, 1793)

Guava Skipper

Panama, Colombia, Venezuela,
Guianas, Trinidad

Phocides distans licinus

(Möschler, 1879)

Distans Skipper

Honduras, Costa Rica, Panama

Phocides metrodorus metrodorus

Bell, 1932

Metrodorus Skipper

Panama to Bolivia and Brazil

Phocides metrodorus nigrescens

Bell, 1938

Nigrescens Skipper

Costa Rica, Colombia, Venezuela,
Ecuador, Peru; species?

Phocides thermus thermus

(Mabille, 1883)

Thermus Skipper

SE Mexico to Venezuela and
Ecuador

Phocides belus

Godman & Salvin, 1890

Belus Skipper

S Texas (one-time stray) to
Costa Rica

Phocides pigmalion pigmalion

(Cramer, 1779)

Pigmalion Skipper

TL: Surinam

Phocides pigmalion okeechobee

(Worthington, 1881)

Mangrove Skipper

S Florida; mangroves

Phocides pigmalion batabanoides

(Holland, 1903)

Mangrove Skipper

most Bahamian Islands

Phocides pigmalion batabano

(Lucas, 1857)

Mangrove Skipper

Cuba, I. de Juventud, Andros,
Little Cayman

Phocides pigmalion bicolora

(Boddaert, 1783)

'Hispaniolan' Mangrove Skipper

Hispaniola

Phocides urania urania

(Westwood, 1852)

Urania Skipper

Mexico, Guatemala; usually in
montane forest

Phocides urania vida

(Butler, 1872)

Vida Skipper

Costa Rica

Phocides lincea perkinsi

(Kaye, 1931)

Perkins' Skipper

Jamaica

Tarsoctenus corytus gaudialis

(Hewitson, 1876)

Gaudy Skipper

Nicaragua to Colombia

Phanus vitreus

(Stoll, 1781)

Widespread Phanus

Oaxaca and Chiapas to southern
Brazil

Phanus ecitonorum

Austin, 1993

Army-ant Phanus

Costa Rica to southern Brazil

Phanus confusis

Austin, 1993

Confusing Phanus

eastern Mexico to El Salvador

Phanus albiapicalis

Austin, 1993

White-tipped Phanus

E & W Mexico to Panama, W
Ecuador and Peru

Phanus rilma

Evans, 1952

West-Mexican Phanus

Pacific coast, Nayarit to
Guatemala

Phanus grandis

Austin, 1993

Grand Phanus

Nicaragua, Costa Rica, Panama,
Colombia, Venezuela

Phanus obscurior obscurior

Kaye, 1925

Dark Phanus

SE Mexico to Colombia and
Trinidad

Phanus marshallii

(Kirby, 1880)

Common Phanus

E & W Mexico to Bolivia

Udranomia orcinus

(C. Felder & R. Felder, 1867)

Orcinus Skipper

SE Mexico to Bolivia

Udranomia kikkawai

(Weeks, 1906)

Nervous Skipper

E & W Mexico to Brazil

Udranomia eurus

(Mabille & Boullet, 1919)

Eurus Skipper

Costa Rica to Venezuela & Amazon
basin

Drephalys helixus

(Hewitson, 1877)

Helixus Skipper

Panama

Drephalys kidonoi

Burns, 1999

Kidono's Skipper

W Costa Rica

Drephalys phoenicoides

(Mabille & Boullet, 1919)

Curious Skipper

Costa Rica to Guianas and NE
Brazil

Drephalys dumeril

(Latreille, [1824])

Dumeril Skipper

Mexico to Ecuador and Brazil

Drephalys oria

Evans, 1952

Oria Skipper

Mexico to Honduras

Drephalys oriander

(Hewitson, 1867)

Oriander Skipper

Mexico, Costa Rica to Guianas
and N Brazil

Drephalys alcmon

(Cramer, 1779)

Alcmon Skipper

Guatemala to Guianas and Brazil

Augiades crinisus

(Cramer, 1780)

Crinisus Skipper

Costa Rica to Bolivia and Brazil

Augiades epimethea bicolor

(Mabille & Boullet, 1919)

Epimethea Skipper

Costa Rica to Bolivia, Guianas
and N Brazil

Hyalothyrus leucomelas

(Geyer, 1832)

Leucomelas Skipper

Panama to Brazil and Peru

Hyalothyrus neleus pemphigargyra

(Mabille, 1888)

Dimorphic Skipper

SE Mexico to Colombia and
Venezuela

Phareas coeleste

Westwood, 1852

Magnificent Skipper

Costa Rica to Bolivia and Brazil

Entheus matho matho

Godman & Salvin, 1879

Giant Entheus

SE Mexico to Costa Rica

Entheus matho aequatorius

Mabille & Boullet, 1919

Equatorial Entheus

Costa Rica to Ecuador

Entheus matho dius

Mabille, 1898

Dius Entheus

Panama Guianas, Peru, and
Bolivia

Entheus crux

Steinhauser, 1989

Mexican Entheus

SE Mexico to Nicaragua

Proteides mercurius mercurius

(Fabricius, 1787)

Mercurial Skipper

continental, S Texas to
Argentina, SE Arizona as rare
stray

Proteides mercurius sanantonio

(Lucas, 1857)

Mercurial Skipper

Cuba, [one-time stray to
Florida]

Proteides mercurius sanchesi

E. Bell & W. Comstock, 1948

Mercurial Skipper

Hispaniola

Proteides mercurius jamaicensis

Skinner, 1920

Mercurial Skipper

Jamaica

Proteides mercurius pedro

(Dewitz, 1877)

Mercurial Skipper

Puerto Rico

Proteides mercurius vincenti

E. Bell & W. Comstock, 1948

Mercurial Skipper

St. Vincent

Proteides mercurius angasi

Godman & Salvin, 1884

Mercurial Skipper

Dominica, St. Lucia, Tobago

Proteides mercurius grenadensis

Enrico & Pinchon, 1969

Mercurial Skipper

Grenada

Proteides maysi

(Lucas, 1857)

Mays' Skipper

Cuba and Isla de Juventud

Epargyreus zestos zestos

(Geyer, 1832)

Zestos Skipper

Florida, Bahamas, Puerto Rico,
Virgin Islands to St. Vincent

Epargyreus zestos inaguarum

Clench & Bjorndal, 1980

Zestos Skipper

Great Inagua, Crooked I., Turks
and Caicos Islands

Epargyreus clarus californicus

MacNeill, 1975

'California' Silver-spotted Skipper

BC to Baja California Norte;
Sierra Nevada-Cascades, westward

Epargyreus clarus

(Cramer, 1775) (Rocky Mountain
segregate)

'Rocky Mountain' Silver-spotted Skipper

NE OR, E WA, ID, MT, WY, C & N
UT, CO, N NM, Black Hills

Epargyreus clarus clarus

(Cramer, 1775)

Silver-spotted Skipper

E USA and Canada to far NE
Mexico (hypothetical)

Epargyreus clarus profugus

Austin, 1998

'Nevada' Silver-spotted Skipper

S NV, N AZ, possibly to central
AZ-NM

Epargyreus clarus huachuca

Dixon, 1955

'Arizona' Silver-spotted Skipper

S AZ, S NM, far NW Mexico

Epargyreus antaeus

(Hewitson, 1867)

Jamaican Silverdrop

Jamaica

Epargyreus spanna

Evans, 1952

Hispaniolan Silverdrop

Hispaniola

Epargyreus socus orizaba

Scudder, 1872

Round-spotted Silverdrop

E Mexico to Guatemala

Epargyreus windi

H. Freeman, 1969

Wind’s Silverdrop

W Mexico

Epargyreus exadeus cruza

Evans, 1952

Broken Silverdrop

(S TX? [CA?]as stray), E & W
Mexico to Panama

Epargyreus aspina

Evans, 1952

Spineless Silverdrop

E & W Mexico to Venezuela and
Ecuador

Epargyreus spina spina

Evans, 1952

Spined Silverdrop

E & W Mexico to Brazil and
Bolivia

Epargyreus spinosa

Evans, 1952

Suffused Silverdrop

W Mexico to Panama

Epargyreus clavicornis tenda

Evans, 1955

Small-spotted Silverdrop

SE Mexico to Venezuela and
Ecuador

Epargyreus clavicornis gaumeri

Godman & Salvin, 1893

Small-spotted Silverdrop

SE Mexico (?) to Honduras

Epargyreus brodkorbi

H. Freeman, 1969

Barely-spotted Silverdrop

Chiapas

Epargyreus deleoni

H. Freeman, 1977

Long-spotted Silverdrop

Yucatan Peninsula (CAMP, QROO,
YUC Mexico) and Belize

Polygonus leo arizonensis

(Skinner, 1911)

Hammock Skipper

SW USA (mostly as strays) & S TX
to Central America

Polygonus leo pallida

Röber, 1925

Hammock Skipper

Central America to Argentina

Polygonus leo histrio

Röber, 1925

Hammock Skipper

Florida and Cuba, some Bahamian
islands

Polygonus leo leo

(Gmelin, 1790)

Hammock Skipper

[= lividus, = ishmael],
Hispaniola, Puerto Rico, SE to
St. Lucia

Polygonus leo hagar

Evans, 1952

Hammock Skipper

Jamaica

Polygonus savigny savigny

(Latreille, [1824])

Manuel’s Skipper

[= manueli], continental, (S TX
as stray), N Mexico to Argentina

Polygonus savigny punctus

E. Bell & W. Comstock, 1948

Manuel’s Skipper

Lesser Antilles; St. Martin S to
Grenadines

Chioides albofasciatus

(Hewitson, 1867)

White-striped Longtail

S TX, S NM, SE AZ, to Costa
Rica; Baja California Sur

Chioides catillus albius

Evans, 1952

Blurry-striped Longtail

Costa Rica and Panama

Chioides churchi

E. Bell & W. Comstock, 1948

Jamaican Longtail

Jamaica

Chioides vintra

Evans, 1952

St. Vincent Longtail

St. Vincent, Grenadines, Grenada

Chioides zilpa

(Butler, 1872)

Zilpa Longtail

S AZ-NM (stray) & S TX, S to
Venezuela and Ecuador

Chioides ixion

(Plötz, 1880)

Hispaniolan Longtail

Hispaniola

Chioides marmorosa

(Herrich-Schäffer, 1865)

Cuban Longtail

Cuba

Aguna asander asander

(Hewitson, 1867)

Gold-spotted Aguna

SE AZ & S TX (as stray) to
Argentina

Aguna asander haitensis

(Mabille & Boullet, 1912)

Gold-spotted Aguna

Cuba and Hispaniola

Aguna asander jasper

Evans, 1952

Gold-spotted Aguna

Jamaica

Aguna claxon

Evans, 1952

Emerald Aguna

Cuba, S Texas to Colombia and
Surinam

Aguna aurunce hypozonius

(Plötz, 1880)

Hewitson's Aguna

SE Mexico to Venezuela

Aguna coelus

(Stoll, 1782)

Stoll's Aguna

Costa Rica and Panama to Peru
and central Brazil

Aguna coeloides

Austin & Mielke, 1998

Austin's Aguna

SE Mexico to southern Brazil

Aguna ganna

(Möschler, 1879)

Ganna Aguna

Costa Rica to Colombia and
Venezuela

Aguna panama

Austin & Mielke, 1998

Panamanian Aguna

Honduras to Venezuela

Aguna metophis

(Latreille, [1824])

Long-tailed Aguna

S Texas to S Brazil

Aguna albistria leucogramma

(Mabille, 1888)

White-striped Aguna

SE Mexico to Colombia and
Venezuela (Surinam?)

Typhedanus undulatus

(Hewitson, 1867)

Mottled Longtail

S Texas (as stray) to Argentina

Typhedanus ampyx

(Godman & Salvin, 1893)

Gold-tufted Skipper

W & NE Mexico to Costa Rica

Typhedanus salas

H. Freeman, 1977

Salas Skipper

S Mexico & Belize

Polythrix octomaculata

(Sepp, [1844])

Eight-spotted Longtail

(SE AZ as stray), S Texas to
Argentina

Polythrix maizae

Hellebuyck, 1998

Melanic Longtail

Nicaragua

Polythrix mexicanus

H. Freeman, 1969

Mexican Longtail

SE AZ, S TX (as stray), N Mexico
to Costa Rica

Polythrix asine

(Hewitson, 1867)

Asine Longtail

N Mexico to Panama

Polythrix caunus

(Herrich-Schäffer, 1869)

Four-spotted Longtail

E Mexico to S Brazil and
Paraguay

Polythrix auginus

(Hewitson, 1867)

Auginus Longtail

SE Mexico to S Brazil

Polythrix metallescens

(Mabille, 1888)

Metallescens Skipper

SE Mexico to Peru and northern
Brazil

Polythrix kanshul

Shuey, 1991

Kanshul Longtail

S Mexico, Belize, NE Costa Rica

Cephise nuspesez

Burns, 1996

Burns' Scarlet-eye

E Mexico to Costa Rica

Cephise mexicanus

Austin & Mielke, 2000

Mexican Scarlet-eye

SLP, OAX (Mexico)

Cephise aelius

(Plötz, 1880)

Longtailed Scarlet-eye

E & S Mexico to Venezuela and
Colombia

Cephise guatemalaensis

(H. Freeman, 1977)

Guatemalan Scarlet-eye

Guatemala, Chiapas, Quintana Roo

Venada advena

(Mabille, 1889)

Advena Scarlet-eye

Panama and Colombia, at least

Venada nevada

Burns, 2005

Nevada Scarlet-eye

Costa Rica

Venada daneva

Burns, 2005

Daneva Scarlet-eye

Costa Rica

Venada cacao

Burns, 2005

Cacao Scarlet-eye

Costa Rica and Panama

Venada naranja

Burns, 2005

Orange Scarlet-eye

Costa Rica

Venada sp. probably undescribed



Cryptic Scarlet-eye

["advena"], Mexico, Guatemala,
(Nicaragua?)

Chrysoplectrum pervivax

(Hübner, [1819])

Pervivax Skipper

Costa Rica to Bolivia, Peru and
N Brazil

Chrysoplectrum perniciosus

(Herrich-Schäffer, 1869)

Perniciosus Skipper

Panama to Bolivia and C Brazil

Chrysoplectrum epicincea

(Butler & H. Druce, 1872)

Stub-tailed Skipper

SE Mexico to Costa Rica

Zestusa dorus

(W. H. Edwards, 1882)

Short-tailed Skipper (Northern Zestusa)

far S CO-UT, AZ, NM, W TX, N
Mexico

Zestusa staudingeri

(Mabille, 1888)

Southern Zestusa

Chiapas to El Salvador

Zestusa elwesi

(Godman & Salvin, 1893)

Mexican Zestusa

Durango to Oaxaca

Zestusa levona

Steinhauser, 1972

Levona's Zestusa

Chiapas to El Salvador

Codatractus carlos carlos

Evans, 1952

Carlos' Mottled-Skipper

C Mexico to Colombia (at least)

Codatractus alcaeus alcaeus

(Hewitson, 1867)

White-crescent Mottled-Skipper

S Texas to Costa Rica

Codatractus alcaeus apulia

Evans, 1952

White-crescent Mottled-Skipper

Panama (?) to W Ecuador

Codatractus yucatanus

H. Freeman, 1977

Yucatan Mottled-Skipper

Yucatan Peninsula

Codatractus arizonensis

(Skinner, 1905)

Arizona Mottled-Skipper

S AZ, S NM, W TX to Oaxaca

Codatractus sallyae

A. Warren, 1995

Sally’s Mottled-Skipper

W Mexico, Yucatan Peninsula,
Guatemala

Codatractus melon

(Godman & Salvin, 1893)

Melon Mottled-Skipper

N Mexico to Costa Rica