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Subfamily Pyrginae Burmeister, 1878   —   Spreadwing Skippers
Tribe Pyrrhopygini Mabille, 1877   —   Firetips, Golf-Club Skippers

Oxynetra hopfferi

Staudinger, 1888

Hopffer´s Firetip

SE Mexico (VER?) to Panama

Azonax typhaon

(Hewitson, 1877)

Typhaon Skipper

SE Mexico (OAX) to Guianas

Myscelus amystis hages

Godman & Salvin, 1893

Widespread Myscelus

E & W Mexico to Panama

Myscelus phoronis phoronis

(Hewitson, 1867)

Phoronis Myscelus

Panama, Colombia, Ecuador, Peru,
Bolivia

Myscelus belti

Godman & Salvin, 1879

Belt's Myscelus

SE Mexico (OAX, VER, CHIS) to
Venezuela

Myscelus perissodora

Dyar, 1914

Dyar's Myscelus

S Mexico (GRO, OAX, VER) to
Colombia

Myscelus assaricus michaeli

Nicolay, 1975

Fiery Myscelus

SE Mexico (OAX, CHIS) to Panama

Passova gellias

(Godman & Salvin, 1893)

Gellias Firetip

Belize, Costa Rica & Panama

Aspitha leander

(Boullet, 1912)

Leander Firetip

SE Mexico (OAX), Colombia

Zonia zonia panamensis

Nicolay, 1975

Pama Skipper

Belize, Panama

Pyrrhopyge zenodorus

Godman & Salvin, 1893

Red-headed Firetip

SE Mexico to Costa Rica

Pyrrhopyge phidias evansi

E. Bell, 1947

Original Firetip

Costa Rica, Panama, Colombia,
Ecuador

Pyrrhopyge pseudophidias

E. Bell, 1931

Impostor Firetip

Panama, Colombia, Ecuador,
Venezuela, Peru, N Brazil

Pyrrhopyge haemon

Godman & Salvin, 1893

Tawny-rimmed Skipper

Costa Rica

Pyrrhopyge crida

Hewitson, 1871

White-banded Firetip

SE Mexico to Colombia and
Ecuador

Creonpyge creon creon

(H. Druce, 1874)

Creon Skipper

N Costa Rica to Cocle Prov.,
Panama, Atlantic & Pacific
slopes

Creonpyge creon lilliana

(Nicolay & Small, 1969)

Creon Skipper

Cerro Campana, far E Cocle, far
W Panama Prov., Panama

Creonpyge creon taylori

(Nicolay & Small, 1981)

Creon Skipper

Fila Cruces, Puntarenas, SE
Costa Rica

Chalypyge chalybea chalybea

(Scudder, 1872)

Orange-rimmed Skipper

central JAL, GTO, QRO

Chalypyge chalybea chloris

(Evans, 1951)

Orange-rimmed Skipper

W and S Mexico, Guatemala,
Honduras

Apyrrothrix araxes arizonae

(Godman & Salvin, 1893)

'Arizona' Araxes Skipper

SE AZ, SW NM, W TX (local), SON,
CHIN, NL, TAMPS, SLP

Apyrrothrix araxes

(Hewitson, 1867)
(Sinaloa-Durango segregate)

Araxes Skipper

SIN, DGO (probably NW NAY)

Apyrrothrix araxes

(Hewitson, 1867) (Nayarit
segregate)

Araxes Skipper

SW NAY

Apyrrothrix araxes

(Hewitson, 1867)
(Guerrero-Michoacan segregate)

Araxes Skipper

GRO, MICH

Apyrrothrix araxes araxes

(Hewitson, 1867)

Araxes Skipper

COL, JAL, DF, MOR, HGO, PUE, VER

Apyrrothrix araxes

(Hewitson, 1867) (Oaxaca
segregate)

Araxes Skipper

OAX

Yanguna cosyra

(H. Druce, 1875)

Cosyra Skipper

Guatemala to Colombia and
Ecuador

Melanopyge mulleri

(E. Bell, 1934)

Red-spotted Skipper

SE Mexico (CHIS, OAX, VER)

Melanopyge erythrosticta

(Godman & Salvin, 1879)

Spotted Skipper

SE Mexico to Panama

Melanopyge sp. undescribed

(Burns)

Dotted Skipper

SE Mexico to Costa Rica

Melanopyge hoffmanni

(H. Freeman, 1977)

Hoffmann's Skipper

SE Mexico (OAX, TAB, CHIS),
Belize

Melanopyge maculosa

(Hewitson, 1866)

Maculosa Skipper

Panama, Colombia, Ecuador

Jonaspyge jonas

(C. Felder & R. Felder, 1859)

Scallop-edged Firetip

SE Mexico (HGO, OAX, PUE, VER,
CHIS), Guat., Nicaragua

Jonaspyge tzotzili

(H. Freeman, 1969)

Freeman's Firetip

SE Mexico (CHIS, OAX)

Jonaspyge aesculapus

(Staudinger, 1876)

Fire-edged Skipper

Costa Rica, Panama, Colombia,
Ecuador

Elbella intersecta strova

Evans, 1951

Variable Firetip

Panama to Venezuela

Elbella patroculus patroculus

(Plötz, 1879)

Patroculus Skipper

Panama to Bolivia

Elbella scylla

(Ménétriés, 1855)

Scylla Firetip

E & W Mexico to Costa Rica

Elbella dulcinea

(Plötz, 1879)

Dulcinea Firetip

Costa Rica, Panama, Colombia,
Venezuela, Ecuador

Elbella patrobas mexicana

Mielke, 1995

Patrobas Skipper

SE Mexico (QROO, TAB, CHIS, VER,
OAX), Belize

Elbella patrobas patrobas

(Hewitson, 1857)

Patrobas Skipper

Guatemala to Colombia

Elbella miodesmiata

(Röber, 1925)

Rober's Skipper

SE Mexico (OAX), Panama,
Colombia

Parelbella macleannani

(Godman & Salvin, 1893)

Macleannan’s Skipper

SE Mexico (CHIS, VER) to Ecuador

Parelbella nigra

Mielke, Austin & A. Warren, 2008

Mielke’s Skipper

SE Mexico (OAX, VER)

Jemadia pseudognetus

(Mabille, 1878)

Dot-collared Skipper

SE Mexico to Venezuela and upper
Amazon

Jemadia pater

Evans, 1951

Pater Skipper

Panama, Colombia, Venezuela

Amysoria galgala

(Hewitson, 1866)

Red-banded Firetip

S Mexico (SW CHIS) to Colombia
and Venezuela

Mysoria barcastus ambigua

(Mabille & Boullet, 1908)

Royal Firetip

SE Mexico (CHIS, TAB, VER) to
Costa Rica

Mysoria barcastus venezuelae

(Scudder, 1872)

Royal Firetip

Panama, Colombia, Venezuela,
Guianas, Trinidad, Tobago

Mysoria amra

(Hewitson, 1871)

Blue-collared Firetip

W Mexico, Guatemala

Mysoria affinis

(Herrich-Schäffer, 1869)

Red-collared Firetip

[= wilsoni], W Mexico, Guatemala

Mysarbia sejanus stolli

Mielke & Casagrande, 2002

Black-veined Firetip

Costa Rica, Colombia, Venezuela,
Fr. Guiana, Upper Amazon
Tribe Celaenorrhinini Swinhoe, 1912   —  

Celaenorrhinus aegiochus

(Hewitson, 1876)

Flashing Flat

Costa Rica and Panama

Celaenorrhinus monartus

(Plötz, 1884)

Dotted Flat

S Mexico to N Brazil

Celaenorrhinus fritzgaertneri

(Bailey, 1880)

Fritzgaertner’s Flat

SE AZ & S TX (as strays) to
Costa Rica

Celaenorrhinus stallingsi

H. Freeman, 1946

Stallings’ Flat

S Texas (rare stray), E Mexico
to Costa Rica

Celaenorrhinus stola

Evans, 1952

Stola Flat

E & W Mexico to Nicaragua

Celaenorrhinus approximatus

R. Williams & E. Bell, 1940

Approximate Flat

Costa Rica and Panama

Celaenorrhinus bifurcus

E. Bell, 1934

Bifurcate Flat

Costa Rica to Guianas, Trinidad,
Ecuador

Celaenorrhinus eligius eligius

(Stoll, 1781)

Eligius Flat

Costa Rica to Guianas, Trinidad,
Tobago, to Peru

Celaenorrhinus cynapes cynapes

(Hewitson, 1870)

Small Flat

SE Mexico to Ecuador
Tribe Carcharodini Verity, 1940   —   Spreadwing Skippers

Mimia phidyle phidyle

(Godman & Salvin, 1894)

Phidyle Skipper

CHIS to Panama

Mimia chiapaensis

H. Freeman, 1969

Chiapan Skipper

CHIS

Arteurotia tractipennis
tractipennis

Butler & H. Druce, 1872

Starred Skipper

S TX to Surinam and Bolivia

Mictris crispus caerula

(Mabille, 1877)

Crisp Skipper

CHIS to Ecuador

Polyctor cleta

Evans, 1953

Cleta Tufted-Skipper

E & W Mexico to Costa Rica

Polyctor polyctor polyctor

(Prittwitz, 1868)

Polyctor Tufted-Skipper

Costa Rica to Argentina

Polyctor enops

(Godman & Salvin, 1894)

Enops Tufted-Skipper

E & S Mexico to Costa Rica

Nisoniades godma

Evans, 1953

Godma Tufted-Skipper

E & S Mexico to Costa Rica

Nisoniades panama

Evans, 1953

Panamanian Tufted-Skipper

Panama to Ecuador

Nisoniades laurentina

(R. Williams & E. Bell, 1939)

Dark Tufted-Skipper

SE Mexico (VER) to Guianas and
Trinidad

Nisoniades rubescens

(Möschler, 1877)

Purplish Tufted-Skipper

W Mexico & S Texas (as stray) to
lower Amazon and Bolivia

Nisoniades ephora

(Herrich-Schäffer, 1870)

Ephora Tufted-Skipper

E & W Mexico to Guianas,
Trinidad, Ecuador and Peru

Nisoniades macarius

(Herrich-Schäffer, 1870)

Macarius Tufted-Skipper

(doubtful for Mexico, Honduras),
Panama to Argentina

Nisoniades castolus

(Hewitson, 1878)

Castolus Tufted-Skipper

SE Mexico (VER) to S Brazil

Nisoniades torta

Steinhauser, 1989

Torta Tufted-Skipper

Costa Rica to Panama

Pachyneuria licisca

(Plötz, 1882)

Immaculate Tufted-Skipper

E & W Mexico to Colombia and
Venezuela

Pellicia arina

Evans, 1953

Glazed Tufted-Skipper

S TX to Panama

Pellicia costimacula costimacula

Herrich-Schäffer, 1870

Frosted Tufted-Skipper

Panama to S Brazil and Paraguay

Pellicia angra angra

Evans, 1953

Rare Tufted-Skipper

SE Mexico (VER) to Panama

Pellicia tonga

Evans, 1953

Tonga Tufted-Skipper

Panama, Colombia, Venezuela

Pellicia dimidiata dimidiata

Herrich-Schäffer, 1870

Morning Glory Tufted-Skipper

W Mexico and S TX (as stray) to
Guianas, Trinidad, up. Amazon

Burca stillmani

E. Bell & W. Comstock, 1948

Stillman's Skipper

Hispaniola

Burca concolor concolor

(Herrich-Schäffer, 1865)

Concolorous Skipper

Cuba

Burca concolor atrata

Rindge, 1955

Concolorous Skipper

Bahamas; Great Bahama Bank

Burca braco braco

(Herrich-Schäffer, 1865)

Braco Skipper

Cuba

Burca braco castigata

Rindge, 1955

Braco Skipper

Bahamas; Great Bahama Bank

Burca cubensis

(Skinner, 1913)

Cuban Skipper

Cuba

Burca hispaniolae

E. Bell & W. Comstock, 1948

Hispaniolan Skipper

Hispaniola

Windia windi

H. Freeman, 1969

Wind’s Skipper

SE Arizona (one-time stray), NW
Mexico to Costa Rica

Noctuana lactifera bipuncta

(Plötz, 1884)

Cryptic Skipper

E & W Mexico to Nicaragua

Noctuana lactifera lactifera

(Butler & H. Druce, 1872)

Cryptic Skipper

Costa Rica to Colombia

Noctuana noctua

(C. Felder & R. Felder, 1867)

Noctua Skipper

Panama to Ecuador and Peru

Noctuana stator

(Godman & Salvin, 1899)

Red-studded Skipper

W Mexico & S TX (one-time stray)
to Guianas, Trinidad, Peru

Morvina fissimacula pelarge

(Godman & Salvin, 1894)

Pelarge Skipper

Nicaragua to Panama

Myrinia myris

(Mabille, 1898)

Myris Skipper

Campeche, Belize, Nicaragua to
Guianas, Bolivia, S Brazil

Myrinia raymundo

H. Freeman, 1979

Raymundo's Skipper

SE Mexico (TAB, CHIS)

Cyclosemia leppa

Evans, 1953

Leppa Eyed-Skipper

Guatemala to Bolivia

Cyclosemia anastomosis

Mabille, 1878

Northern Eyed-Skipper

E Mexico to Ecuador

Cyclosemia lyrcaea

(Hewitson, 1878)

Lyrcaea Eyed-Skipper

Panama to SE Brazil

Cyclosemia pedro

R. Williams & E. Bell, 1940

Pedro Eyed-Skipper

Honduras to N Brazil ("Amazons")

Cyclosemia caerulea

Austin & DeVries, 2001

Caerulea Eyed-Skipper

Costa Rica

Cyclosemia subcaerulea

Schaus, 1913

Subcaerulea Eyed-Skipper

Costa Rica to Ecuador

Gorgopas chlorocephala
chlorocephala

(Herrich-Schäffer, 1870)

Green-headed Sootywing

Nicaragua to Bolivia

Bolla cylindus

(Godman & Salvin, 1896)

Checkered Sootywing

SE Mexico (VER) to Costa Rica

Bolla cybele

Evans, 1953

Veracruzan Sootywing

SE Mexico (VER)

Bolla cyclops

(Mabille, 1876)

Cyclops Sootywing

[= sonda], SE Mexico, Guatemala,
Colombia

Bolla atahuallpai

(Lindsey, 1925)

Atahuallpai Sootywing

Panama to Argentina

Bolla cupreiceps

(Mabille, 1891)

Copper-headed Sootywing

E & S Mexico to Bolivia and S
Brazil

Bolla subapicatus

(Schaus, 1902)

Fin-tipped Sootywing

W & S Mexico, Guatemala

Bolla orsines

(Godman & Salvin, 1896)

Hook-tipped Sootywing

E & W Mexico to El Salvador

Bolla evippe

(Godman & Salvin, 1896)

Rough-tipped Sootywing

E & W Mexico, Guatemala,
Nicaragua

Bolla imbras

(Godman & Salvin, 1896)

Rounded Sootywing

[= pullata], E & S Mexico to
Colombia and Venezuela

Bolla brennus brennus

(Godman & Salvin, 1896)

Obscure Sootywing

S Texas (one-time stray), E
Mexico to Panama

Bolla oriza

Evans, 1953

Orizaba Sootywing

SE Mexico (VER, CHIS?)

Bolla guerra

Evans, 1953

Guerrero Sootywing

SW Mexico to Nicaragua

Bolla boliviensis

(E. Bell, 1937)

Bolivian Sootywing

Guatemala to Bolivia

Bolla saletas

(Godman & Salvin, 1896)

Saletas Sootywing

Guatemala to El Salvador

Bolla eusebius

(Plötz, 1884)

Spatulate Sootywing

E & W Mexico to Costa Rica

Bolla fenestra

Steinhauser, 1991

Oaxacan Sootywing

S Mexico (OAX)

Bolla solitaria

Steinhauser, 1991

Solitary Sootywing

S Mexico (OAX)

Bolla zorilla

(Plötz, 1886)

Zorilla Sootywing

SE Mexico (VER, OAX) to Bolivia

Bolla clytius

(Godman & Salvin, 1897)

Mottled Sootywing

SE AZ & S TX (rare strays) to
Honduras

Bolla litus

(Dyar, 1912)

Many-spotted Sootywing

mostly W Mexico

Staphylus ceos

(W. H. Edwards, 1882)

Golden-headed Sootywing

far S CA, S NV (rare), S AZ, S
NM, S TX, to QRO

Staphylus vulgata

(Möschler, 1879)

Golden-snouted Sootywing

E & W Mexico to Argentina

Staphylus caribbea

(R. Williams & E. Bell, 1940)

Panamanian Sootywing

Nicaragua to Ecuador

Staphylus tepeca

(E. Bell, 1942)

Grizzled Sootywing

C & S Mexico (mostly DF to OAX)

Staphylus hayhurstii

(W. H. Edwards, 1870)

Hayhurst's Sootywing

S Ontario, E USA to far SE SD,
NB, KS, to S-C TX

Staphylus tierra

Evans, 1953

West-Mexican Sootywing

W Mexico (SON to CHIS), El
Salvador

Staphylus mazans

(Reakirt, [1867])

Mazans Sootywing

C TX to CHIS

Staphylus ascalaphus

(Staudinger, 1876)

Central American Sootywing

SE Mexico to Bolivia, S Brazil

Staphylus perforata

(Möschler, 1879)

Perforata Sootywing

Panama, Colombia, Peru

Staphylus unicornis

Steinhauser & Austin, 1993

Unicorn Sootywing

Costa Rica

Staphylus lenis

Steinhauser, 1989

Lenis Sootywing

Yucatan Peninsula to Trinidad

Staphylus azteca

(Scudder, 1872)

Aztec Sootywing

E & W Mexico to Costa Rica

Staphylus vincula

(Plötz, 1886)

Mountain Sootywing

E & W Mexico to Venezuela

Staphylus oeta

(Plötz, 1884)

Oeta Sootywing

Costa Rica to Bolivia and
Argentina

Staphylus parvus

Steinhauser & Austin, 1993

Parvus Sootywing

Costa Rica

Staphylus evemerus

Godman & Salvin, 1896

Evemerus Sootywing

Nicaragua to Panama

Staphylus huigra huigra

(R. Williams & E. Bell, 1940)

Huigra Sootywing

El Salvador, Ecuador to Bolivia

Staphylus iguala

(R. Williams & E. Bell, 1940)

Iguala Sootywing

W & S Mexico

Staphylus cartagoa

(R. Williams & E. Bell, 1940)

Cartagoa Sootywing

Costa Rica, El Salvador

Staphylus esmeraldus

L. Miller, 1966

Esmeraldus Sootywing

E Costa Rica

Pholisora catullus

(Fabricius, 1793)

Common Sootywing

widespread in USA and S Canada,
S to Colima and Hidalgo...S to
Oaxaca

Pholisora mejicanus

(Reakirt, [1867])

Mexican Sootywing

S-C Colorado to Oaxaca

Hesperopsis alpheus oricus

(W. H. Edwards, 1879)

Saltbush Sootywing

E & S CA, NV, SW UT, NW AZ