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Texola elada ulrica (W. H. Edwards, 1877)
(Elada Checkerspot)

Pinned Specimens photo collection:


Live Adults photo collection Page 1:

Live Adults photo collection Page 2:

Immatures photo collection:

Larval Foodplants and Habitats photo collection:

Distribution and Larval Foodplants:

C & W TX, NE Mexico

Larval Foodplants: Acanthaceae: Carlowrightia parviflora (Buckl.) Wasshausen, Siphonoglossa pilosella (Nees) Torr.


(From: A Catalogue of the Butterflies of the United States and Canada, Jonathan P. Pelham)

Genus Texola Higgins, 1959

Lepid. News 12(5-6): 161, figs. 1b, 2b ♂ genitalia, 3b palp, 4b FW veination (dymas), 5b FW (dymas). Type-species: Eresia elada Hewitson, 1868, Ill. exot. Butts. 4(33): [49], pl. Eresia VII [19], figs. 54, 55, by original designation.


Texola elada (Hewitson, 1868)

Ill. exot. Butts. 4(33): [49], pl. Eresia VII [19], figs. 54 ♂ D, 55 ♂ V.

Original Combination: Eresia Elada

Type Locality: “Mexico”

Types: Holotype in BMNH.

            =    callina (Boisduval, 1869)

Annls. Soc. Entomol. Belg. 12: 54, no. 39.

Original Combination: Melitaea Callina

Type Locality: “dans la Sonora, nous avons reçu du Mexique plusiers individus de cette même espece”; lectotype is labeled, probably erroneously, “Sonora.”

Types: Lectotype in CMNH, designated by Higgins (1960), Trans. R. Entomol. Soc. Lond. 112(14): 452-453; figured by W. Holland (1931), Butt. Book (rev. ed.): pl. 59, fig. 21 ♂ D. F. Brown (1965), J. Lepid. Soc. 19(4): fig. 4 ♂ D&V, and J. Emmel et al. (1998), Syst. W. N. Am. Butts. (2): 31, figs. 231 ♂ D, 232 ♂ V.

J. Emmel et al. (1998), ibid., discussed this taxon and the uncertainty about its type locality. The lectotype was loaned to W. H. Edwards by Boisduval, but Boisduval died before it could be returned. The subsequent figure of a specimen of Texola elada ulrica by Oberthür (1914), Étud. Lépid. Comp. 9(2): pl. 260, fig. 2185 ♂ D&V, and designation of that figure as the “type” of callina by W. Barnes and McDunnough (1916), Contr. Nat. Hist. Lepid. N. Am. 3(2): 92, confused the issue; F. Brown (1965), J. Lepid. Soc. 19(4): 207-211, detailed this confusion. The actual type locality remains unknown; there currently are no populations of Texola in Lorquin’s “Sonora” (the vicinity of Los Angeles, California).

            =    socia (R. Felder, 1869)

Verh. zool.-bot. Ges. Wien 19: 470-471, no. 18.

Original Combination: Eresia socia

Type Locality: not stated; “Mexico” implied from the title of the paper.

Types: Location of type(s) not known.


Texola elada ulrica (W. H. Edwards, 1877)

Can. Entomol. 9(10): 189-190.

Original Combination: Melitaea ulrica

Type Locality: “San Antonio, Texas” [Bexar County]

Types: Lectotype in CMNH, designated by Higgins (1960), Trans. R. Entomol. Soc. Lond. 112(14): 454; figured by W. Holland (1931), Butt. Book (rev. ed.): 132, pl. 17, fig. 2 ♂ D, and by F. Brown (1966), Trans. Am. Entomol. Soc. 92(3): 383, fig. 9 ♂ D&V.

                   =    imitata (Strecker, [1878])

Lepid. Rhop. Het. (14): 130.

Original Combination: Melitaea Imitata

Type Locality: Not specifically stated; “vicinity of New Braunfels and San Antonio, Southwestern Texas” implied from introductory text. [Comal and Bexar counties]

Types: Syntypes in FMNH.


Brown, Frederick Martin. 1944. Notes on Mexican Butterflies, IV. Journal of the New York entomological Society 52(4): 343-360. {[19] Dec 1944} 

Barnes, William and James Halliday McDunnough. 1916. Notes on North American diurnal Lepidoptera. Contributions to the Natural History of the Lepidoptera of North America 3(2): 49-156, 8 pls. {15 Dec 1916} 

Boisduval, Jean Baptiste Alphonse d'Echauffour. 1868-1869. Lépidoptères de la Californie. Annales de la Société entomologique de Belgique 12: 1-28 (1868); 37-94 ([1 Nov] 1869) [missing pp. 29-37 through printer's error] {1868-1869} [description of Melitaea Callina

Brown, Frederick Martin. 1965. Three letters from J. A. B. D. De Boisduval to W. H. Edwards, and the true identity of Melitaea pola Bdv. and Melitaea callina Bdv. Journal of the Lepidopterists= Society 19(4): 197-211, 5 figs. {20 Dec 1965} 

Brown, Frederick Martin. 1966. The types of the nymphalid butterflies described by William Henry Edwards--Part II, Melitaeinae. Transactions of the American Entomological Society 92(3): 357-468, 34 figs. {Sep, 20 Oct 1966} 

Brown, Frederick Martin. 1967. Lorquin's localities "Sonora" and "Utah." Journal of the Lepidopterists= Society 21(4): 271-274. {28 Nov 1967} 

Edwards, William Henry. 1877. Descriptions of new species of butterflies belonging to the N. American Fauna. Canadian Entomologist 9(10): 189-192. {[31] Oct 1877} [description of Melitaea ulrica

Emmel, John F., Thomas Chadbourne Emmel and Sterling O. Mattoon. 1998. 2. The types of California butterflies named by Jean Alphonse Boisduval: designation of lectotypes and a neotype, and fixation of type localities. Pp. 3-76, 269 figs., in: T. C. Emmel (Ed.), Systematics of western North American butterflies. Gainesville; Mariposa Press; {[18] Dec 1998} 

Godman, Frederick du Cane and Osbert Salvin. 1879-1901. Biologia Centrali-Americana. Zoology: Insecta. Lepidoptera-Rhopalocera. London; Taylor & Francis, 3 volumes (Vols. 36-38): 1(Nym., Rio.)(1879-1886); xlv+487 pp.; 2(Pier., Pap., Lyc., Hes.)(1887-1901): 782 pp.; 3(plates): 1-48 (1879-1886); 49-112 (1887-1901) 

Hewitson, William Chapman. 1852-1877. Illustrations of new species of exotic butterflies, selected chiefly from the collections of W. Wilson Saunders and William C. Hewitson. London; John Van Voorst, 4to, 5 vols., 100 parts; pages unnumbered; 300 pls. [description of Eresia Elada]

Higgins, Lionel George. 1959. Four new melitaeine genera (Nymphalidae). Lepidopterists= News 12(5-6): 161-164, 5 figs. {1958, 30 Apr 1959} [description of Texola

Higgins, Lionel George. 1960. A revision of the melitaeine genus Chlosyne and allied species (Lepidoptera: Nymphalidae). Transactions of the Royal Entomological Society of London 112(14): 381-467, 134 figs. {31 Dec 1960} 

Hoffmann, Carlos Christian. 1940. Catálogo systemático y zoogeográphico de los Lepidópteros Mexicanos, Primera parte. Papilionoidea. Anales del Instituto de Biología. Universidad Nacional de México 11(2): 639-739, 2 maps. {[after 28 Dec] 1940} 

Holland, William Jacob. 1931. The butterfly book. New and thoroughly revised edition.  A popular and scientific manual, describing and depicting all the butterflies of the United States and Canada. Garden City, New York; Doubleday, Doran & Co.  xii + 424 pp., 77 pls., 198 + [1] text figures. {1931}

Kendall, Roy O=Neal. 1960. More larval foodplants from Texas. Journal of the Lepidopterists= Society 13(4): 221-228 {1959, 1 Aug 1960} 

Lamas, Gerardo. 1999. Nomenclatural faux‑pas among western North American butterflies. Lepidoptera News 1999(2): 7-8 {30 June [19 Aug] 1999} 

Maza, Roberto F. de la. 1987. Mariposas Mexicanas. Guía para su colecta y determinación. México; Fondo de Cultura Económica: 304 pp., 67 pls., 58 figs. {Oct 1987} [Ed. 2 1991] 

Röber, Johannes Karl Max 1913-1914. Gattung Phyciodes. In: A. Seitz (Ed.), Die Gross-Schmetterlinge der Erde 5(162): pl. 92 {5 Jun 1913}; (183): 434-440, pl. 90 {10 Dec 1913}; (188): 449-450, pls. 88, 89. {[before 10 Feb] 1914}; (189): pl. 91. {14 Apr 1914} 

Strecker, Ferdinand Heinrich Herman. 1874. 14. New Lep. from the vicinity of New Braunfels and San Antonio, Texas. Pp. 130-132, in: Lepidoptera, rhopaloceres and heteroceres, indigenous and exotic; with descriptions and colored illustrations. Reading, Pennsylvania; Private. {Sep 1877, [18 Mar 1878]} [description of Melitaea Imitata]

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